Yva Momatiuk & John Eastcott / Waterfall and peak, Fitzroy Massif, Los Glaciares National Park, Patagonia, Argentina

Yva Momatiuk & John Eastcott / Waterfall and peak, Fitzroy Massif, Los Glaciares National Park, Patagonia, Argentina

Alex Mustard / Veined octopus (Amphioctopus marginatus) sheltering in an old clam shell, with plankton swirling around it at night (captured as trails in the long exposure). Bitung, North Sulawesi, Indonesia. Lembeh...

Hiroya Minakuchi / Japanese Firefly (Luciola cruciata) group flying along river at night, Shikoku, Japan

Alex Mustard / Yellow tube sponge (Aplysina fistularis) on a coral reef, with the camera rotated during the long exposure. East End, Grand Cayman, Cayman Islands. British West Indies. Caribbean Sea.

SCOTLAND: The Big Picture / View across Sound of Taransay to North Harris hills in stormy weather , West Harris, Outer Hebrides, Scotland, UK, September 2014.

Alex Mustard / Boulder brain coral (Colpophyllia natans) photographed with a long exposure with camera rotation. Taken with circular fisheye lens. Grand Cayman, Cayman Islands. Caribbean Sea.

Bence Mate / Flock of Great egrets (Ardea alba) at twilight, Lake Csaj, Kiskunsagi National Park, Hungary. May.

Bence Mate / Black crowned night heron (Nycticorax nycticorax) at twilight with other herons in the background, including Grey heron (Ardea cinerea) in flight, Lake Csaj, Kiskunsagi National Park, Pusztaszer, Hun...

Pete Oxford / Kaieteur Falls, Potaro River, Kaieteur National Park, Guyana

Chien Lee / Stream in lowland rainforest, Masoala National Park, Antsiranana, Madagascar

Matthias Breiter / Trees in autumn along river, Sand River, Agawa Bay, Ontario, Canada

Matthias Breiter / Creek flowing into lake in temperate rainforest, Lake Crescent, Olympic National Park, Washington

Matthias Breiter / Waterfall and creek in temperate rainforest, Merriman Falls, Olympic National Park, Washington

Benjamin Barthelemy / Long exposure of tidal water flowing off rocks, with the Okshornan Mountains in the background, Isle of Senja, Troms, Norway, February 2015.

Wim van den Heever / Quiver tree (Aloe dichotoma) with the Milky Way at night, and light pollution from town in the distance, Keetmanshoop, Namibia. Colours accentuated digitally.

Juan Carlos Munoz / Water cascading down Toberia falls, Andoin, Sierra Entzia Natural Park, Alava, Basque Country, Spain, April.

Ben Hall / Marine iguana (Amblyrhynchus cristatus) on rock taken with slow shutter speed to show motion in the water, Galapagos islands, May.

Wim van den Heever / Nile crocodile (Crocodylus niloticus) still in the fast flowing water of the Mara River, taken with slow shutter speed. Masai Mara, Kenya.

Jack Dykinga / Lightning storm with rainbow over Saguaro cactus (Carnegiea gigantea) Avra Valley, Saguaro National Park, Tucson Mountains, Arizona, USA, August 2015. Long exposure with lightning trigger.

Jack Dykinga / Lightning storm at twilight with Saguaro cactus (Carnegiea gigantea) Avra Valley, Saguaro National Park, Tucson Mountains, Arizona, USA. June 2015. Long exposure with lightning trigger.

Bence Mate / Wild boar (Sus scrofa) at night with the milky way in the background, Gyulaj, Tolna, Hungary. August. Taken using long exposure with flash at night. Winner of the Mammals Category of the GDT Awards 20...

Mark MacEwen / Japanese firefly (Luciola cruciata) light trails at night above stream in rural Japan. June.

David Tipling / Northern gannet (Sula bassana) colony, Hermaness, Unst, Shetland Scotland, UK, April.

Ross Hoddinott / Trevose Lighthouse at dusk, long exposure at high tide, Trevose, Cornwall, UK. January 2015.

Ross Hoddinott / Ice on black volcanic sand beach with waves, Jokulsarlon, Southeast Iceland, February.

Peter Cairns / Selfoss waterfall in autumn with snow, north Iceland, September 2013.

Peter Cairns / View over fishing village of Hamnoya, Lofoten, Norway. March 2015.

Sean Crane / Bat Star (Asterina miniata) trio at low tide, Montana De Oro State Park, Los Osos, California

Yukihiro Fukuda / Light trails of Japanese fireflies (Hotaria parvula and Luciola cruciata), Nichinan-chou, Tottori, Japan, July, Endemic species.

Ole Jorgen Liodden / Chatham stream on the main Chatham island, New Zealand, March 2014.

Colin Monteath / Coastal cliffs, Tunnel Beach, Otago Peninsula, Otago, New Zealand

Alex Hyde / Long exposure of light trail from bioluminescent click beetle (Pyrophorus sp) walking on ground at night. Danum Valley Sabah, Borneo.

Alex Hyde / Takob-Akob Falls plunging 38 metres through the rainforest. Southern plateau edge, Maliau Basin, Sabah, Borneo, May 2011.

Floris van Breugel / Mount Fitz Roy at dawn on a snowy winter morning, Patagonia region of Argentina near El Chalten, June 2014.

Floris van Breugel / Salto Grande waterfall with Paine Grande beyond, Torres del Paine National Park, Chile, June 2014.

Christophe Courteau / Storm at 'Pointe de Pern', Ile d'Ouessant / Ushant. Nividic lighthouse visible in distance. Finistere, Brittany, France, September 2011.

Luciano Candisani / Jacare Caiman (Caiman yacare) group in wetland at night, Pantanal, Brazil

David Noton / Stars and Milky Way over Durdle Door and the Jurassic Coast, with the lights of Weymouth and Portland beyond, Dorset, England, UK, December 2013.

Sven Zacek / Roe deer (Capreolus capreolus) running through snow, Vorumaa, Estonia, January. Winner of the Animal stories portfolio in the Melvita Nature Images Awards competition 2014.

Sam Hobson / Fallow deer (Dama dama) herd walking down road. London, UK. January.

Sven Zacek / Ural owl (Strix uralensis) flying to nest with star trails in sky above, Estonia, May. Winner of the Birds category in the Melvita Nature Images Awards competition 2014.

Hougaard Malan / Aloes below the snow-capped peaks of the Swartberg. Near Calitzdorp, Western Cape, South Africa. June 2009.

Hougaard Malan / Lighthouse at sunset. Kommetjie, Cape Town, South Africa. November 2008.

Hougaard Malan / Waterfall and autumn foliage with view to Mount Fitz Roy. El Chalten, Patagonia, Argentina. April 2013.

Sam Hobson / Rose-ringed / ring-necked parakeets (Psittacula krameri) in flight on way to roost in an urban cemetery, London, UK, January. Finalist in the Birds category, Wildlife Photographer of the Year Awards (...

Bence Mate / Greenfinches (Carduelis chloris) and House sparrows (Passer domesticus) blurred long exposure photograph, Hungary, February.

Ingo Arndt / Sunrise at Iguacu Falls, largest waterfalls in the world, Iguacu National Park, Brazil

Alex Mustard / A long exposure renders Bar jacks (Carangoides ruber) as ghostly trails as they swim around a sponge, on a coral reef, at night. East End, Grand Cayman, Cayman Islands. Overall winner of GDT competiti...

Nature Production / Japanese firefly (Luciola cruciata) on leaf at night, with light trails of hundreds of other fireflies in flight in the background, Japan

Yukihiro Fukuda / Japanese fireflies (Luciola cruciata) in flight at night, Japan endemic species, Hino-River, Nichinan-chou, Tottori, Japan, July

Yukihiro Fukuda,Yukihiro Fukuda / Japanese fireflies (Luciola cruciata) in flight at night, endemic species, Yaku-shima UNESCO World Heritage Site, Kagoshima, Japan, May

Inaki Relanzon / Long exposure of Godafoss waterfall, Iceland.

Ben Lascelles / Nightime long exposure of the Zhong Chu river below Zutruphuk Monastery, Mount Kailash. Tibet. June 2010

Tim Fitzharris / Ruby Beach, Olympic National Park, Washington

Adrian Davies / Long exposure of a Sensitive plant (Mimosa pudica), showing movement of leaves and stem following irritation.

Mark Hamblin / 2020VISION / Receding wave shot with long exposure. Talisker Bay, Isle of Skye, Scotland, UK, October 2011.

Peter Cairns / 2020VISION / Gannet (Morus bassanus) breeding colony, long exposure showing flight trails. Hermaness National Nature Reserve, Shetland, Scotland, July.

Eric Medard / Natterer's Bat (Myotis nattereri) flying low over water to drink, with long-exposure star-trails. France, Europe, July.

Jack Dykinga / Cypress dome swamp with water surface covered with water-spangles and duckweed, Big Cypress Seminole Indian Reservation, Florida Everglades, USA

Konrad Wothe / Three Snow Geese (Chen caerulescens atlanticus / Chen caerulescens) in flight, Bosque del Apache, New Mexico, USA, November

Peter Cairns / 2020VISION / Long exposure of sea, with rocks in foreground, Bagh Dhail Mor, Isle of Lewis, Outer Hebrides, Scotland, UK, October 2011

Sven Zacek / Northern lights far on the horizon in night landscape. Sarek National Park, Sweden, Europe, September.

Mark Hamblin / 2020VISION / Hen harrier (Circus cyaneus) adult female in flight, moorland habitat, long exposure, Glen Tanar Estate, Grampian, Scotland, UK, June

Peter Cairns / 2020VISION / Stormy seas off Hosta, North Uist, Western Isles / Outer Hebrides, Scotland, UK, May 2011

Andy Rouse / 2020VISION / Dipper (Cinclus cinclus) perched on rock in river, long exposure, Brecon Beacons NP, Wales, UK, May

Bence Mate / Hoopoe (Upupa epops) in flight, Hungary. Multi flash, long exposure. Winner, Eric Hosking award, 2011 Wildlife Photographer of the Year competition.

Juergen Freund / Lunar tailed rock cod / Coronation grouper (Variola louti) swimming fast over coral reef, Bali, Indonesia

Inaki Relanzon / Dolmen d'Eyna at night, with a long exposure capturing star trails. Eyne, Pyrenees, France, September 2010.

Peter Cairns / 2020VISION / View over Gannet colony (Morus bassanus) with flight trails of birds in flight, Hermaness NNR, Unst, Shetland, Scotland, June 2010

Ben Hall / 2020VISION / Birch trees blowing in high winds, long exposure, Calke Abbey, The National Forest, Central England, UK, November 2010

Chris Gomersall / 2020VISION / Flock of Red knot (Calidris canutus) in flight, long exposure, winter plumage, Snettisham RSPB reserve, Norfolk, England, UK, October. Did you know? The Wash Estuary in Norfolk is estimated to have a...

Chris Gomersall / 2020VISION / Roosting flock of Red knot (Calidris canutus) long exposure, winter plumage, Snettisham RSPB reserve, Norfolk, England, UK, October. 2020VISION Book Plate.

Chris Gomersall / 2020VISION / Roosting flock of Red knot (Calidris canutus) long exposure, winter plumage, Snettisham RSPB reserve, Norfolk, England, UK, October

Inaki Relanzon / Star trails above Baobab trees (Adansonia za) at night, Zombitse National Park, Madagascar, June 2010

Luciano Candisani / Waterfall, Chapada dos Veadeiros National Park, Goias State, Brazil

Chien Lee / Waterfalls draining remote highland plateau, Lawas, Sarawak, Borneo, Malaysia

Colin Monteath / Waterfall below Mount Fitzroy, Los Glaciares National Park, Patagonia, Argentina

Colin Monteath / Waterfall at dawn below Mount Fitzroy, Los Glaciares National Park, Patagonia, Argentina

Jerry Monkman / Sunrise near Brenton Point State Park, from Ocean Road, Newport, Rhode Island, USA, October 2009

Floris van Breugel / Long exposure to capture star trails circling around the North Star, Polaris, at the "Racetrack," a remote desert playa (dry lake bed) in Death Valley National Park, famous for its mysterious moving r...

Alex Mustard / Long exposure image of Tiger shark (Galeocerdo cuvier) chasing bait, Bahamas.

Edwin Giesbers / Sunlight falling on Bluebells (Hyacinthoides non-scripta / Endymion scriptum) flowering in Beech wood, Hallerbos, Belgium, April, long exposure

Edwin Giesbers / Bluebells (Hyacinthoides non-scripta / Endymion scriptum) flowering in Beech wood, Hallerbos, Belgium, April, long exposure

Sebastian Kennerknecht / Coastline at dusk, Shark Fin Cove, Davenport, California

Chien Lee / Small waterfall in rainforest interior, Lambir Hills National Park, Borneo, Malaysia

Laurie Campbell / Long exposure of wave breaking on rocks at sunset, Caithness, Scotland, August

Jack Dykinga / Virgin river flowing through Zion Canyon Narrows, with reflections of afternoon sun, Zion National Park, Utah, USA

Wild Wonders of Europe / Zankl / Waves lapping against rocks on the Dwejra coast, Gozo, Malta, May 2009

Jeff Vanuga / Grizzly bear {Ursus arctos horribilis} beside water, long exposure image, Yellowstone National Park, Wyoming, USA, October

Wild Wonders of Europe / Popp-Ha / Matterhorn (4,478m) at night, long exposure with star trails, viewed from Gornergrat, Switzerland, September 2008, climbers in foreground with headlamps setting off to climb the Matterhorn from Hornli...

Jerry Monkman / Moores Brook, Ellsworth, Maine, USA. Moores Brook empties into Branch Lake. June 2008

Stephen Belcher / Anak Krakatau erupting in May 2009, Ujung Kulon National Park, Sunda Strait, Indonesia

Ross Hoddinott / Porth Nanvan SSSI (Cot Valley) looking towards the Brisons, Cornwall, UK. Evening light. November 2008.

Ross Hoddinott / St Michael's Mount at sunrise, from Marazion beach, Cornwall, UK. November 2008.

Kevin Schafer / Iguacu Falls, world's largest waterfalls, Brazil and Argentina border