Books - Secrets of the Seas (155) More files...
With photographs by Alex Mustard and text by Callum Roberts, Secrets of the Seas, published in September 2016 by Bloomsbury, is a visual voyage to some of the richest and most exciting places in the world's oceans. With chapters covering a range of habitats from the Pacific's Coral Triangle to the North Atlantic and the ocean's seaweed cathedrals, the book showcases Alex Mustard's extraordinary photography, with remarkable close-ups of a diverse range of the ocean's inhabitants.

“This isn’t a title to read just once, but one into which you’ll continue to dive – to escape, to be entertained, and to become better informed about our fascinating, spectacular oceans.” – BBC Wildlife
Grey seal (Halichoerus grypus) portrait underwater, Farne Islands, Northumberland, England, UK, August
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Alex Mustard / 2020VISION / Grey seal (Halichoerus grypus) portrait underwater, Farne Islands, Northumberland, England, UK, August
School of Bohar snappers (Lutjanus bohar) above a coral reef. These fish are usually solitary but have gathered in the summer in the Red Sea for spawning. Shark Reef, Ras Mohammed Marine Park, Sinai,...
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Alex Mustard / School of Bohar snappers (Lutjanus bohar) above a coral reef. These fish are usually solitary but have gathered in the summer in the Red Sea for spawning. Shark Reef, Ras Mohammed Marine Park, Sinai,...
Schoolmaster snapper (Lutjanus apodus) school, under a coral ledge, Snapper Hole, East End, Grand Cayman, Cayman Islands, Caribbean Sea.
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Alex Mustard / Schoolmaster snapper (Lutjanus apodus) school, under a coral ledge, Snapper Hole, East End, Grand Cayman, Cayman Islands, Caribbean Sea.
Portrait of Whale shark (Rhincodon typus) at the surface. Isla Mujeres, Quintana Roo, Yucatan Peninsular, Mexico. Caribbean Sea.
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Alex Mustard / Portrait of Whale shark (Rhincodon typus) at the surface. Isla Mujeres, Quintana Roo, Yucatan Peninsular, Mexico. Caribbean Sea.
Great hammerhead shark (Sphyrna mokarran) portrait, South Bimini, Bahamas. The Bahamas National Shark Sanctuary. Gulf Stream, West Atlantic Ocean.
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Alex Mustard / Great hammerhead shark (Sphyrna mokarran) portrait, South Bimini, Bahamas. The Bahamas National Shark Sanctuary. Gulf Stream, West Atlantic Ocean.
Harlequin swimming crab (Lissocarcinus laevis) sheltering beneath the tentacles of a tube anemone (Cerianthus sp.). Dauin, Dumaguete, Negros, Philippines. Bohol Sea, Tropical West Pacific Ocean.
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Alex Mustard / Harlequin swimming crab (Lissocarcinus laevis) sheltering beneath the tentacles of a tube anemone (Cerianthus sp.). Dauin, Dumaguete, Negros, Philippines. Bohol Sea, Tropical West Pacific Ocean.
Caribbean reef shark (Carcharhinus perezi) bursts through the middle of a school of Yellowtail snappers (Ocyurus chrysurus). Grand Bahama, Bahamas. Tropical West Atlantic Ocean.
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Alex Mustard / Caribbean reef shark (Carcharhinus perezi) bursts through the middle of a school of Yellowtail snappers (Ocyurus chrysurus). Grand Bahama, Bahamas. Tropical West Atlantic Ocean.
Fangtooth moray eel (Enchelycore anatina) with mouth open, Grand Canaria, Canary Islands, Spain. East Atlantic Ocean.
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Alex Mustard / Fangtooth moray eel (Enchelycore anatina) with mouth open, Grand Canaria, Canary Islands, Spain. East Atlantic Ocean.
Secretary blenny (Acanthemblemari maria) yawns as it peers out from a hole in the reef. West Bay, Grand Cayman, Cayman Islands, British West Indies. Caribbean Sea.
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Alex Mustard / Secretary blenny (Acanthemblemari maria) yawns as it peers out from a hole in the reef. West Bay, Grand Cayman, Cayman Islands, British West Indies. Caribbean Sea.
Nassau grouper (Epinephelus striatus) yawning on a coral reef drop off. San Salvador island, Bahamas. Bahamas Sea, Tropical West Atlantic Ocean.
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Alex Mustard / Nassau grouper (Epinephelus striatus) yawning on a coral reef drop off. San Salvador island, Bahamas. Bahamas Sea, Tropical West Atlantic Ocean.
Leafy seadragon (Phycodurus eques). Wool Bay Jetty, Edithburgh, Yorke Peninsula, South Australia.
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Alex Mustard / Leafy seadragon (Phycodurus eques). Wool Bay Jetty, Edithburgh, Yorke Peninsula, South Australia.
Spotlight goby (Gobiosoma louisae) looking out from inside a Yellow tube sponge (Aplysina fistularis), growing on a coral reef. Bloody Bay Wall, Little Cayman, Cayman Islands, British West Indies. Car...
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Alex Mustard / Spotlight goby (Gobiosoma louisae) looking out from inside a Yellow tube sponge (Aplysina fistularis), growing on a coral reef. Bloody Bay Wall, Little Cayman, Cayman Islands, British West Indies. Car...
Close up detail photo of a Spotted moray (Gymnothorax isingteena), Baa Atoll, Maldives. Indian Ocean.
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Alex Mustard / Close up detail photo of a Spotted moray (Gymnothorax isingteena), Baa Atoll, Maldives. Indian Ocean.
Large female Southern stingray (Hypanus americanus) in shallow water, under storm clouds, Stingray City, Grand Cayman. Stingray City is feeding site for stingrays and bait was used on this shoot. Caym...
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Alex Mustard / Large female Southern stingray (Hypanus americanus) in shallow water, under storm clouds, Stingray City, Grand Cayman. Stingray City is feeding site for stingrays and bait was used on this shoot. Caym...
Pattern detail of a Majestic angelfish (Pomacanthus navarchus). Fiabacet Islands, Misool, Raja Ampat, West Papua, Indonesia. Ceram Sea.
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Alex Mustard / Pattern detail of a Majestic angelfish (Pomacanthus navarchus). Fiabacet Islands, Misool, Raja Ampat, West Papua, Indonesia. Ceram Sea.
Pink shrimp (Pandalus montagui), Loch Carron, Ross and Cromarty, Scotland, UK, April
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Alex Mustard / 2020VISION / Pink shrimp (Pandalus montagui), Loch Carron, Ross and Cromarty, Scotland, UK, April
An Australian Sea Lion (Neophoca cinerea) bathing in the sun at the surface. Kangaroo Island, South Australia, Australia. Southern Ocean. November
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Alex Mustard / An Australian Sea Lion (Neophoca cinerea) bathing in the sun at the surface. Kangaroo Island, South Australia, Australia. Southern Ocean. November
Whale shark (Rhincodon typus) silhouette, feeding on floating fish eggs (not visible) just below the surface in calm weather. Note the bow wave ahead of the shark. Isla Mujeres, Quintana Roo, Yucatan...
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Alex Mustard / Whale shark (Rhincodon typus) silhouette, feeding on floating fish eggs (not visible) just below the surface in calm weather. Note the bow wave ahead of the shark. Isla Mujeres, Quintana Roo, Yucatan...
California sea lion (Zalophus californianus) adult female swimming through sunbeams in an underwater cave. Los Isotes, La Paz, Mexico. Sea of Cortez, East Pacific Ocean.
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Alex Mustard / California sea lion (Zalophus californianus) adult female swimming through sunbeams in an underwater cave. Los Isotes, La Paz, Mexico. Sea of Cortez, East Pacific Ocean.
Yellow lipped sea snake (Laticauda colubrina) dives back down to the reef after surfacing for breath of air. Apo Island, Dumaguete, Negros, Philippines. Bohol Sea, Tropical West Pacific Ocean.
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Alex Mustard / Yellow lipped sea snake (Laticauda colubrina) dives back down to the reef after surfacing for breath of air. Apo Island, Dumaguete, Negros, Philippines. Bohol Sea, Tropical West Pacific Ocean.
Harlequin shrimp (Hymenocera elegans) on a blue sea star. This species of shrimps feeds on starfish, turning them upside-down and keeping them as a living larder. Bitung, North Sulawesi, Indonesia. Le...
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Alex Mustard / Harlequin shrimp (Hymenocera elegans) on a blue sea star. This species of shrimps feeds on starfish, turning them upside-down and keeping them as a living larder. Bitung, North Sulawesi, Indonesia. Le...
Basking shark (Cetorhinus maximus) with mouth wide open feeding on plankton concentrated in surface waters close to the island of Coll, Inner Hebrides, Scotland, UK, June 2011. 2020VISION Exhibition....
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Alex Mustard / 2020VISION / Basking shark (Cetorhinus maximus) with mouth wide open feeding on plankton concentrated in surface waters close to the island of Coll, Inner Hebrides, Scotland, UK, June 2011. 2020VISION Exhibition....
Great white shark (Carcharodon carcharias) from above. Guadalupe Island, Baja California, Mexico. East Pacific Ocean
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Alex Mustard / Great white shark (Carcharodon carcharias) from above. Guadalupe Island, Baja California, Mexico. East Pacific Ocean
Air filled bladders of Giant kelp (Macrocystis pyrifera). Santa Barbara Island, Channel Islands. Los Angeles, California, United States of America. North East Pacific Ocean.
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Alex Mustard / Air filled bladders of Giant kelp (Macrocystis pyrifera). Santa Barbara Island, Channel Islands. Los Angeles, California, United States of America. North East Pacific Ocean.
Boulder brain coral (Colpophyllia natans) photographed with a long exposure with camera rotation. Taken with circular fisheye lens. Grand Cayman, Cayman Islands. Caribbean Sea.
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Alex Mustard / Boulder brain coral (Colpophyllia natans) photographed with a long exposure with camera rotation. Taken with circular fisheye lens. Grand Cayman, Cayman Islands. Caribbean Sea.
Nudibranchs (Doto greenamyeri) newly described species on feather hydroids. Seraya Bay, north coast of Bali, Indonesia. Java Sea.
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Alex Mustard / Nudibranchs (Doto greenamyeri) newly described species on feather hydroids. Seraya Bay, north coast of Bali, Indonesia. Java Sea.
Pacific giant octopus (Enteroctopus dofleini) hunts for food on a colourful reef in Browning Pass, Vancouver Island, British Columbia, Canada
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Alex Mustard / Pacific giant octopus (Enteroctopus dofleini) hunts for food on a colourful reef in Browning Pass, Vancouver Island, British Columbia, Canada
Harbour seal (Phoca vitulina) in the kelp forest (Macrocystis pyrifera). Santa Barbara Island, Channel Islands. Los Angeles, California, United States of America. North East Pacific Ocean.
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Alex Mustard / Harbour seal (Phoca vitulina) in the kelp forest (Macrocystis pyrifera). Santa Barbara Island, Channel Islands. Los Angeles, California, United States of America. North East Pacific Ocean.
Pink skeleton shrimp (Caprella sp.) portrait in red algae. Thistilfjord, Thorshofn, north east Iceland. North Atlantic Ocean.
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Alex Mustard / Pink skeleton shrimp (Caprella sp.) portrait in red algae. Thistilfjord, Thorshofn, north east Iceland. North Atlantic Ocean.
RF- Pair of Rock beauties (Holacanthus tricolor) spawning at dusk. The larger male rubs against the flank of female during the spawning rise. Cayman Islands, British West Indies. Caribbean Sea. (This...
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Alex Mustard / RF- Pair of Rock beauties (Holacanthus tricolor) spawning at dusk. The larger male rubs against the flank of female during the spawning rise. Cayman Islands, British West Indies. Caribbean Sea. (This...
Aggregation of tiny isopods (Santia sp.) living on the surface of a Blue sponge (Haliclona sp.), there are two colour varieties here, possibly different species. There are also small copepods living o...
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Alex Mustard / Aggregation of tiny isopods (Santia sp.) living on the surface of a Blue sponge (Haliclona sp.), there are two colour varieties here, possibly different species. There are also small copepods living o...
Brandt's cormorant (Phalacrocorax penicillatus) bursts through a school of Pacific chub mackerel (Scomber japonicus) while hunting beneath an oil rig. Ellen Rig, Los Angeles, California, United States...
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Alex Mustard / Brandt's cormorant (Phalacrocorax penicillatus) bursts through a school of Pacific chub mackerel (Scomber japonicus) while hunting beneath an oil rig. Ellen Rig, Los Angeles, California, United States...
Diver swimming with an aggregation of Cod (Gadus morhua) over a kelp forest. These cod were gathered in early spring off the north coast of Iceland to spawn. Thorshofn, Iceland. North Atlantic Ocean....
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Alex Mustard / Diver swimming with an aggregation of Cod (Gadus morhua) over a kelp forest. These cod were gathered in early spring off the north coast of Iceland to spawn. Thorshofn, Iceland. North Atlantic Ocean....
A school of lesser devil rays (Mobula hypostoma) flying through sunbeams as they feed. Isla Mujeres, Quintana Roo, Yucatan Peninsula, Mexico. Caribbean Sea.
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Alex Mustard / A school of lesser devil rays (Mobula hypostoma) flying through sunbeams as they feed. Isla Mujeres, Quintana Roo, Yucatan Peninsula, Mexico. Caribbean Sea.
Silhouette of reef fish, including a Blue triggerfish (Pseudobalistes fuscus) and many Scalefin anthias (Pseudanthias squamipinnis). Ras Mohammed Marine Park, Sinai, Egypt. Red Sea.
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Alex Mustard / Silhouette of reef fish, including a Blue triggerfish (Pseudobalistes fuscus) and many Scalefin anthias (Pseudanthias squamipinnis). Ras Mohammed Marine Park, Sinai, Egypt. Red Sea.
Soft corals (Scleronephthya corymbosa) inside a cavern within a coral reef. This species is typical of cave habitats. Uum Karerim, St Johns Reef. Egypt. Red Sea.
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Alex Mustard / Soft corals (Scleronephthya corymbosa) inside a cavern within a coral reef. This species is typical of cave habitats. Uum Karerim, St Johns Reef. Egypt. Red Sea.
A male Weedy Seadragon (Phyllopteryx taeniolatus) carrying eggs along tail swims through a Kelp (Macrocystis pyrifera) forest  in Tasmania, Australia. Tasmania is the only part of Australia with giant...
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Alex Mustard / A male Weedy Seadragon (Phyllopteryx taeniolatus) carrying eggs along tail swims through a Kelp (Macrocystis pyrifera) forest in Tasmania, Australia. Tasmania is the only part of Australia with giant...
Diver photographing a school of Big eye trevally (Caranx sexfasciatus). South Atoll, Tubbataha Atolls, Tubbataha Reefs Natural Park, Palawan, Philippines. Sulu Sea.
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Alex Mustard / Diver photographing a school of Big eye trevally (Caranx sexfasciatus). South Atoll, Tubbataha Atolls, Tubbataha Reefs Natural Park, Palawan, Philippines. Sulu Sea.
Colourful reef scene with Green tubastrea (Tubastrea micrantha), Soft corals (Scleronephthya sp.) and Scalefin anthias (Pseudanthias squamipinnis) and Magenta slender anthias (Luzonichthys waitei). Ra...
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Alex Mustard / Colourful reef scene with Green tubastrea (Tubastrea micrantha), Soft corals (Scleronephthya sp.) and Scalefin anthias (Pseudanthias squamipinnis) and Magenta slender anthias (Luzonichthys waitei). Ra...
Aggregation of Golden jellyfish (Mastigias sp.) in a marine lake in Palau, the golden colour of this species comes from symbiotic algae in its tissues. Jellyfish Lake, Eil Malk island, Rock Islands, P...
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Alex Mustard / Aggregation of Golden jellyfish (Mastigias sp.) in a marine lake in Palau, the golden colour of this species comes from symbiotic algae in its tissues. Jellyfish Lake, Eil Malk island, Rock Islands, P...
Sponge crab (Dromia dromia) carrying a purple sponge on its back, has climbed on a pinky-orange sponge to feed at night. Baa Atoll, Maldives.
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Alex Mustard / Sponge crab (Dromia dromia) carrying a purple sponge on its back, has climbed on a pinky-orange sponge to feed at night. Baa Atoll, Maldives.
Diver encounters a large sunfish (Mola mola), the world's heaviest bony fish. Tulamben, Bali, Indonesia. Java Sea ~Model released
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Alex Mustard / Diver encounters a large sunfish (Mola mola), the world's heaviest bony fish. Tulamben, Bali, Indonesia. Java Sea ~Model released
Close up of the scale details of a Yellow-mask angelfish (Pomacanthus xanthometopon). Baa Atoll, Maldives. Indian Ocean
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Alex Mustard / Close up of the scale details of a Yellow-mask angelfish (Pomacanthus xanthometopon). Baa Atoll, Maldives. Indian Ocean
Lionfish (Pterois volitans) portrait, invasive species in the Caribbean. East End, Grand Cayman, Cayman Islands, British West Indies, Caribbean Sea.
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Alex Mustard / Lionfish (Pterois volitans) portrait, invasive species in the Caribbean. East End, Grand Cayman, Cayman Islands, British West Indies, Caribbean Sea.
Florida manatee (Trichechus manatus latirostrus) being cleaned by blue gill sunfish (Lepomis macrochirus). Three Sisters Spring, Crystal River, Florida. USA. February 2010
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Alex Mustard / Florida manatee (Trichechus manatus latirostrus) being cleaned by blue gill sunfish (Lepomis macrochirus). Three Sisters Spring, Crystal River, Florida. USA. February 2010
Four California spiny lobsters (Panulirus interruptus) shelter beneath a boulder in a kelp forest. Santa Barbara Island, Channel Islands. Los Angeles, California, United States of America. North East...
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Alex Mustard / Four California spiny lobsters (Panulirus interruptus) shelter beneath a boulder in a kelp forest. Santa Barbara Island, Channel Islands. Los Angeles, California, United States of America. North East...
School of Blacksmith fish (Chromis punctipinnis) swim through a giant kelp (Macrocystis pyrifera) forest. Santa Barbara Island, Channel Islands. Los Angeles, California, United States of America. Nort...
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Alex Mustard / School of Blacksmith fish (Chromis punctipinnis) swim through a giant kelp (Macrocystis pyrifera) forest. Santa Barbara Island, Channel Islands. Los Angeles, California, United States of America. Nort...
Snorkeller swimming over a shallow coral reef with a large stand of Staghorn coral (Acropora cervicornis). North Wall, Grand Cayman, Cayman Islands, British West Indies. Caribbean Sea. Critically enda...
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Alex Mustard / Snorkeller swimming over a shallow coral reef with a large stand of Staghorn coral (Acropora cervicornis). North Wall, Grand Cayman, Cayman Islands, British West Indies. Caribbean Sea. Critically enda...
School of Silversides (Atherinomorus lacunosus) mass below a jetty, creating a false ceiling of fish. Berenice Jetty, Aqaba, Jordan. Gulf of Aqaba, Jordan.
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Alex Mustard / School of Silversides (Atherinomorus lacunosus) mass below a jetty, creating a false ceiling of fish. Berenice Jetty, Aqaba, Jordan. Gulf of Aqaba, Jordan.
Bumphead parrotfish (Bolbometopon muricatum) gather at dawn on a shallow coral reef. Sipadan Island, Sabah, Borneo, Malaysia. Celebes Sea. Vulnerable species.
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Alex Mustard / Bumphead parrotfish (Bolbometopon muricatum) gather at dawn on a shallow coral reef. Sipadan Island, Sabah, Borneo, Malaysia. Celebes Sea. Vulnerable species.
Rich coral growth, with a pair of Exquiste butterflyfish (Chaetodon austriacus), flourishes beneath the barren desert cliffs in the Red Sea. Taken at sunset. Ras Katy, Sinai, Egypt. Gulf of Aqaba, Red...
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Alex Mustard / Rich coral growth, with a pair of Exquiste butterflyfish (Chaetodon austriacus), flourishes beneath the barren desert cliffs in the Red Sea. Taken at sunset. Ras Katy, Sinai, Egypt. Gulf of Aqaba, Red...
Great white sharks (Carcharodon carcharias) males beneath the surface. Guadalupe Island, Baja California, Mexico. East Pacific Ocean
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Alex Mustard / Great white sharks (Carcharodon carcharias) males beneath the surface. Guadalupe Island, Baja California, Mexico. East Pacific Ocean
Red gorgonians (Lophogorgia chilensis) growing beneath a Giant kelp (Macrocystis pyrifera) forest, with female California sheephead (Semicossyphus pulcher). Santa Barbara Island, Channel Islands. Los...
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Alex Mustard / Red gorgonians (Lophogorgia chilensis) growing beneath a Giant kelp (Macrocystis pyrifera) forest, with female California sheephead (Semicossyphus pulcher). Santa Barbara Island, Channel Islands. Los...
Southern stingrays (Hypanus americanus) swimming over a sand bar in the early morning. Grand Cayman, Cayman Islands. British West Indies. Caribbean Sea.
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Alex Mustard / Southern stingrays (Hypanus americanus) swimming over a sand bar in the early morning. Grand Cayman, Cayman Islands. British West Indies. Caribbean Sea.
Portrait of a Green sea turtle (Chelonia mydas) in morning light. Sipadan Island, Sabah, Borneo, Malaysia. Celebes Sea.
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Alex Mustard / Portrait of a Green sea turtle (Chelonia mydas) in morning light. Sipadan Island, Sabah, Borneo, Malaysia. Celebes Sea.
Diver swimming past a huge coral head. Buyat Bay, North Sulawesi, Indonesia. Lembeh Strait, Molucca Sea.
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Alex Mustard / Diver swimming past a huge coral head. Buyat Bay, North Sulawesi, Indonesia. Lembeh Strait, Molucca Sea.
Underwater photographer Alex Mustard taking a self-portrait with a friendly young grey seal (Halichoerus grypus). Farne Islands, Northumberland, England, British Isles. North Sea. ~ Model released.
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Alex Mustard / Underwater photographer Alex Mustard taking a self-portrait with a friendly young grey seal (Halichoerus grypus). Farne Islands, Northumberland, England, British Isles. North Sea. ~ Model released.
California sea lion (Zalophus californianus) resting in the canopy of a forest of giant kelp (Macrocystis pyrifera). Santa Barbara Island, Channel Islands. Los Angeles, California, United States of Am...
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Alex Mustard / California sea lion (Zalophus californianus) resting in the canopy of a forest of giant kelp (Macrocystis pyrifera). Santa Barbara Island, Channel Islands. Los Angeles, California, United States of Am...
Red-spotted guard crab (Trapezia tigrina) female on a (Pocillopora sp.) coral. Seraya, Tulamben, Bali, Indonesia. Java Sea
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Alex Mustard / Red-spotted guard crab (Trapezia tigrina) female on a (Pocillopora sp.) coral. Seraya, Tulamben, Bali, Indonesia. Java Sea
Caribbean reef shark (Carcharhinus perezi) swimming over Common sea fans (Gorgonia ventalina) on a coral reef. East End, Grand Cayman, Cayman Islands, British West Indies. Caribbean Sea
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Alex Mustard / Caribbean reef shark (Carcharhinus perezi) swimming over Common sea fans (Gorgonia ventalina) on a coral reef. East End, Grand Cayman, Cayman Islands, British West Indies. Caribbean Sea
Bohar snappers (Lutjanus bohar) breaking up into smaller groups to spawn. The fish swim short arches above the main group as they release clouds of gametes. ~Shark City, Ulong, Rock Islands, Palau. Tr...
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Alex Mustard / Bohar snappers (Lutjanus bohar) breaking up into smaller groups to spawn. The fish swim short arches above the main group as they release clouds of gametes. ~Shark City, Ulong, Rock Islands, Palau. Tr...
Amphipod travelling on sea gooseberry (Ctenophora) Browning Pass, Vancouver Island, British Columbia, Canada. North East Pacific Ocean.
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Alex Mustard / Amphipod travelling on sea gooseberry (Ctenophora) Browning Pass, Vancouver Island, British Columbia, Canada. North East Pacific Ocean.
Long exposure of a juvenile Harlequin sweetlips (Plectorhinchus chaetodonoides), which dance as a defence from predators. The dance may mimic the movements of poisonous flatworms or may simply confuse...
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Alex Mustard / Long exposure of a juvenile Harlequin sweetlips (Plectorhinchus chaetodonoides), which dance as a defence from predators. The dance may mimic the movements of poisonous flatworms or may simply confuse...
Brandt's cormorant (Phalacrocorax penicillatus) swimming beneath an oil rig. Eureka Rig, Los Angeles, California, United States of America. North East Pacific Ocean.
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Alex Mustard / Brandt's cormorant (Phalacrocorax penicillatus) swimming beneath an oil rig. Eureka Rig, Los Angeles, California, United States of America. North East Pacific Ocean.
California sea lions (Zalophus californianus) group playing in the sun in the early morning. Santa Barbara Island, Channel Islands. Los Angeles, California, United States of America. North East Pacifi...
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Alex Mustard / California sea lions (Zalophus californianus) group playing in the sun in the early morning. Santa Barbara Island, Channel Islands. Los Angeles, California, United States of America. North East Pacifi...
Portrait of a male Squarespot anthias (Pseudanthias pleurotaenia). Sangeang Island; Sumbawa, Indonesia. Flores Sea.
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Alex Mustard / Portrait of a male Squarespot anthias (Pseudanthias pleurotaenia). Sangeang Island; Sumbawa, Indonesia. Flores Sea.
Portrait of a Nudibranch (Flabellina pellucida) on a kelp frond with star trails. Gulen, Norway. North East Atlantic Ocean.
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Alex Mustard / Portrait of a Nudibranch (Flabellina pellucida) on a kelp frond with star trails. Gulen, Norway. North East Atlantic Ocean.
Mimic octopus (Thaumoctopus mimicus) mating/courtship. The smaller male is riding on top of the female as he tries to place his sperm sac inside her mantle with his arm. Lembeh Strait, Sulawesi, Indon...
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Alex Mustard / Mimic octopus (Thaumoctopus mimicus) mating/courtship. The smaller male is riding on top of the female as he tries to place his sperm sac inside her mantle with his arm. Lembeh Strait, Sulawesi, Indon...
California sea lions (Zalophus californianus) babies playing in the safety of shallow water.  Santa Barbara Island, Channel Islands. Los Angeles, California, United States of America. North East Pacif...
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Alex Mustard / California sea lions (Zalophus californianus) babies playing in the safety of shallow water. Santa Barbara Island, Channel Islands. Los Angeles, California, United States of America. North East Pacif...
Portrait of a male Stocky anthias (Pseudanthias hypselosoma). Seraya, Tulamben, Bali, Indonesia. Java Sea
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Alex Mustard / Portrait of a male Stocky anthias (Pseudanthias hypselosoma). Seraya, Tulamben, Bali, Indonesia. Java Sea
Nudibranch (Polycera quadrilineata) on kelp fronds. Gulen, Norway. North East Atlantic Ocean.
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Alex Mustard / Nudibranch (Polycera quadrilineata) on kelp fronds. Gulen, Norway. North East Atlantic Ocean.
Portrait of a male River blenny (Salaria fluviatilis) River Flumendosa, Gennargentu National Park, Sardinia, Italy.
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Alex Mustard / Portrait of a male River blenny (Salaria fluviatilis) River Flumendosa, Gennargentu National Park, Sardinia, Italy.
Harbour seal (Phoca vitulina) investigating the leg of the photographer (Alex Mustard). Santa Barbara Island, Channel Islands. Los Angeles, California, United States of America. North East Pacific Oce...
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Alex Mustard / Harbour seal (Phoca vitulina) investigating the leg of the photographer (Alex Mustard). Santa Barbara Island, Channel Islands. Los Angeles, California, United States of America. North East Pacific Oce...
Starfish close up detail, Blairgowrie, Mornington Penisular, Victoria, Australia. Port Philip Bay, Bass Strait.
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Alex Mustard / Starfish close up detail, Blairgowrie, Mornington Penisular, Victoria, Australia. Port Philip Bay, Bass Strait.
Cleaner wrasse (Labroides dimidiatus) swims out from the gill of a Blue spotted pufferfish (Arothron caeruleopunctatus). East of Eden, Similan Islands, Thailand. Andaman Sea, Indian Ocean.
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Alex Mustard / Cleaner wrasse (Labroides dimidiatus) swims out from the gill of a Blue spotted pufferfish (Arothron caeruleopunctatus). East of Eden, Similan Islands, Thailand. Andaman Sea, Indian Ocean.
Detail of pattern of an adult Emperor angelfish (Pomacanthus imperator) Seraya, Tulamben, Bali, Indonesia. Java Sea.
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Alex Mustard / Detail of pattern of an adult Emperor angelfish (Pomacanthus imperator) Seraya, Tulamben, Bali, Indonesia. Java Sea.
A portrait of a small Candy stripe shrimp (Lebbeus grandimanus) in front of its host pink anemone (out of focus because of the magnification). Browning Wall, Browning Pass, Port Hardy, Vancouver Islan...
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Alex Mustard / A portrait of a small Candy stripe shrimp (Lebbeus grandimanus) in front of its host pink anemone (out of focus because of the magnification). Browning Wall, Browning Pass, Port Hardy, Vancouver Islan...
Slender grouper (Anyperodon leucogrammicus) exploiting the confusion caused by and Orange spotted trevally (Carangoides bajad) hunting silversides (Atherinidae) to plan an ambush. Raja Ampat, West Pap...
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Alex Mustard / Slender grouper (Anyperodon leucogrammicus) exploiting the confusion caused by and Orange spotted trevally (Carangoides bajad) hunting silversides (Atherinidae) to plan an ambush. Raja Ampat, West Pap...
Studio image of multiple Nudibranch species. Gulen, Norway. North East Atlantic Ocean. Digital composite.
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Alex Mustard / Studio image of multiple Nudibranch species. Gulen, Norway. North East Atlantic Ocean. Digital composite.
Abstract view of Seafan (Melithaea sp.). Ra Province, Viti Levu, Fiji. Nananu Passage, Vatu-i-Ra Passage, Bligh Waters, Tropical South Pacific Ocean.
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Alex Mustard / Abstract view of Seafan (Melithaea sp.). Ra Province, Viti Levu, Fiji. Nananu Passage, Vatu-i-Ra Passage, Bligh Waters, Tropical South Pacific Ocean.
Imitator damselfish (Pomacentrus imitator) male, abstract in in front of a Black coral tree (Antipathes sp.) on a coral reef. Ra Province, Viti Levu, Fiji, Polynesia. Bligh Waters, Vatu-i-Ra Passage,...
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Alex Mustard / Imitator damselfish (Pomacentrus imitator) male, abstract in in front of a Black coral tree (Antipathes sp.) on a coral reef. Ra Province, Viti Levu, Fiji, Polynesia. Bligh Waters, Vatu-i-Ra Passage,...
Hairy frogfish (Antennarius striatus) on a sandy seabed. Anilao, Batangas, Luzon, Philippines. Verde Island Passages, Tropical West Pacific Ocean.
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Alex Mustard / Hairy frogfish (Antennarius striatus) on a sandy seabed. Anilao, Batangas, Luzon, Philippines. Verde Island Passages, Tropical West Pacific Ocean.
Split level photo of group of Sockeye salmon (Oncorhynchus nerka) fighting their way upstream as they migrate back to the river of their birth to spawn, trees showing autumnal colours. Adams River, Br...
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Alex Mustard / Split level photo of group of Sockeye salmon (Oncorhynchus nerka) fighting their way upstream as they migrate back to the river of their birth to spawn, trees showing autumnal colours. Adams River, Br...
Coral grouper (Cephalopholis miniata) guarding its territory on a colourful coral reef. East Of Eden, Similan Islands, Thailand. Andaman Sea, Indian Ocean.
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Alex Mustard / Coral grouper (Cephalopholis miniata) guarding its territory on a colourful coral reef. East Of Eden, Similan Islands, Thailand. Andaman Sea, Indian Ocean.
Steller sealion (Eumetopias jubatus) male charging past the camera. Race Rocks, Victoria, Vancouver Island, British Columbia. Canada. North East Pacific Ocean.
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Alex Mustard / Steller sealion (Eumetopias jubatus) male charging past the camera. Race Rocks, Victoria, Vancouver Island, British Columbia. Canada. North East Pacific Ocean.
Dome coral (Porites nodifera) pinnacle or bommie surrounded by fish, including Scalefin anthias (Pseudanthias squamipinnis), with diver Fury Shoal, Egypt. Red Sea.
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Alex Mustard / Dome coral (Porites nodifera) pinnacle or bommie surrounded by fish, including Scalefin anthias (Pseudanthias squamipinnis), with diver Fury Shoal, Egypt. Red Sea.
Longfin batfish (Platax teira) with Fusiliers (Caesio spp.) in the current in front of Baby Rock pinnacle. Misool, Raja Ampat, West Papua, Indonesia. Ceram Sea.
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Alex Mustard / Longfin batfish (Platax teira) with Fusiliers (Caesio spp.) in the current in front of Baby Rock pinnacle. Misool, Raja Ampat, West Papua, Indonesia. Ceram Sea.
Lion's mane jellyfish (Cyanea capillata) with feeding tentacles spread swimming in the sun in shallow water. Gulen, Norway. North East Atlantic Ocean.
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Alex Mustard / Lion's mane jellyfish (Cyanea capillata) with feeding tentacles spread swimming in the sun in shallow water. Gulen, Norway. North East Atlantic Ocean.
Butterfish (Pholis gunnellus) portrait. Gardur, south west Iceland. North Atlantic Ocean.
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Alex Mustard / Butterfish (Pholis gunnellus) portrait. Gardur, south west Iceland. North Atlantic Ocean.
Veined octopus (Amphioctopus marginatus) shelters in discarded coconut shell. Lembeh Strait, North Sulawesi, Indonesia. Molucca Sea.
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Alex Mustard,Alex Mustard / Veined octopus (Amphioctopus marginatus) shelters in discarded coconut shell. Lembeh Strait, North Sulawesi, Indonesia. Molucca Sea.
Blue shark (Prionace glauca) swimming near the surface of the English Channel. Penzance, Cornwall, England, British Isles. North East Atlantic Ocean, August. Highly commended in the Portraits category...
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Alex Mustard / Blue shark (Prionace glauca) swimming near the surface of the English Channel. Penzance, Cornwall, England, British Isles. North East Atlantic Ocean, August. Highly commended in the Portraits category...
Blue sharks (Prionace glauca) cruise beneath the surface of the English Channel. Penzance, Cornwall, England, British Isles. North East Atlantic Ocean, August. Winner of the Coast and Marine category...
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Alex Mustard / Blue sharks (Prionace glauca) cruise beneath the surface of the English Channel. Penzance, Cornwall, England, British Isles. North East Atlantic Ocean, August. Winner of the Coast and Marine category...
A large colony of Mountainous star coral (Montastraea faveolata) spawning at night on a coral reef, showing the synchronous release of bundles of eggs and sperm, East End, Grand Cayman, Cayman Islands...
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Alex Mustard / A large colony of Mountainous star coral (Montastraea faveolata) spawning at night on a coral reef, showing the synchronous release of bundles of eggs and sperm, East End, Grand Cayman, Cayman Islands...
Rear view of a Basking shark (Cetorhinus maximus), Cairns of Coll, Inner Hebrides, Scotland, UK, June.
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Alex Mustard / 2020VISION / Rear view of a Basking shark (Cetorhinus maximus), Cairns of Coll, Inner Hebrides, Scotland, UK, June.
A close up portrait of a cod (Gadus morhua) over a kelp forest. Husavik Iceland. North Atlantic Ocean. April 2012
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Alex Mustard / A close up portrait of a cod (Gadus morhua) over a kelp forest. Husavik Iceland. North Atlantic Ocean. April 2012
Young Grey seal (Halichoerus grypus) exploring a seaweed garden in summer, Cairns Of Coll, Island of Coll, Inner Hebrides, Scotland, UK, North Atlantic Ocean, June. Overall Winner of the Marine Catego...
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Alex Mustard / 2020VISION / Young Grey seal (Halichoerus grypus) exploring a seaweed garden in summer, Cairns Of Coll, Island of Coll, Inner Hebrides, Scotland, UK, North Atlantic Ocean, June. Overall Winner of the Marine Catego...