Christian Ziegler / Orchid (Trigonidium egertonianum) flower, central Panama

Edwin Giesbers / Saltwater crocodile (Crocodylus porosus) close up portrait, Sarawak, Borneo, Malaysia

Laurent Geslin / Urban Red fox {Vulpes vulpes} + angry domestic cat in church graveyard, London, UK

Cyril Ruoso / Drill (Mandrillus leucophaeus) adult male appeasement face, Pandrillus Drill Sanctuary, Nigeria

Cyril Ruoso / Mountain Gorilla (Gorilla gorilla beringei) large silverback continuing to display after attacking a tourist, Virunga Mountains, Rwanda

Alex Mustard / Bluestriped grunt (Haemulon sciurus) portrait, West End, Grand Cayman, Cayman Islands, British West Indies, Caribbean Sea.

Andy Rouse / Polar Bear (Ursus maritimus) in snowstorm. Spitsbergen, Norway. Non-excusive.

Daniel Heuclin / Cocks fighting in cock fight, Guimaras island, Philippines, February 2007

Juergen Freund / Longfin Bannerfish (Heniochus acuminatus) head on portrait, Bali, Indonesia, Indo-pacific

Juergen Freund / Sargassum frogfish / anglerfish (Histrio histrio) in its floating Sargassum seaweed home, El Nido, Palawan, Philippines

Fabio Liverani / Freshwater Crab (Potamon fluviatile) brandishing claws in defensive posture. Foreste Casentinesi National Park, Italy, July.

Alex Mustard / Lemon shark (Negaprion brevirostris) in shallow water with reflection at the surface. Little Bahama Bank, Bahamas.

Petra Wegner / Alaskan Malamute portrait of blue coated adult.

Anup Shah / African lion (Panthera leo) adult male fighting back against the adolescent male who has disturbed his sleep, Masai Mara National Reserve, Kenya. March.

Stephen Dalton / Harlequin beetle (Acrocinus longimanus) in flight, Venezuela, South America.

Edwin Giesbers / Head portrait of Hamadryas baboon (Papio hamadryas) Captive. Apenheul zoo; the Netherlands.

David Shale / Angler fish Melanocetus murrayi, Mid-Atlantic Ridge, North Atlantic Ocean

Jose B. Ruiz / Portrait of Iberian wolf {Canis lupus sygnatus} snarling, captive, Lobo Park, Antequera, Malaga, Spain

Pete Oxford / Red-legged Seriema (Cariama cristata) threat display, Brazil

Michael Durham / Amur Leopard (Panthera pardus orientalis) portrait, native to the Amur River Valley, Siberia, Manchuria and Korea

ZSSD / Western Lowland Gorilla (Gorilla gorilla gorilla) male vocalizing, native to equatorial West Africa

Konrad Wothe / Mountain Gorilla (Gorilla gorilla beringei) males fighting, Virunga National Park, Democratic Republic of the Congo

Matthias Breiter / Polar Bear (Ursus maritimus) adult male charging opponent, Churchill, Manitoba, Canada

Gerry Ellis / Western Lowland Gorilla (Gorilla gorilla gorilla) silverback male, equatorial Africa

Matthias Breiter / Kodiak Bear (Ursus arctos middendorffi) with mouth agape, native to Kodiak Island, Alaska

Konrad Wothe / Grizzly Bear (Ursus arctos horribilis) growling, North America

Lynn M Stone / Melanistic (black form) leopard snarling, often called black panther (captive)

Anup Shah / Face portrait of snarling Bengal tiger {Panthera tigris tigris} India

Donald M. Jones / Lesser Scaup (Aythya affinis) male, central Montana

Mark Taylor / Collie cross dogs Brec and Bliss, exchanging angry snarls. Brec with hackles raised.

Thomas Marent / Gelada Baboon (Theropithecus gelada) male in defensive display, Simien Mountains National Park, Ethiopia

Thomas Marent / Brown Capuchin (Cebus apella), Manu National Park, Peru

Thomas Marent / Bonobo (Pan paniscus) in defensive posture, Lola Ya Bonobo Sanctuary, Democratic Republic of the Congo

Thomas Marent / Buff-cheeked Gibbon (Nomascus gabriellae) female in defensive posture, Phnom Tamao Wildlife Rescue Center, Cambodia

Sean Crane / Burrowing Owl (Athene cunicularia) perching on a termite mound, Piaui State, Brazil

Winfried Wisniewski / Golden Jackal (Canis aureus), Keoladeo Nationa Park, India

Winfried Wisniewski / Golden Jackal (Canis aureus), Keoladeo Nationa Park, India

Thomas Marent / Golden Snub-nosed Monkey (Rhinopithecus roxellana) male in threat display, Qinling Mountains, Shaanxi, China

Cyril Ruoso / Drill (Mandrillus leucophaeus) adult male appeasement face, Pandrillus Drill Sanctuary, Nigeria

Cyril Ruoso / Mountain Gorilla (Gorilla gorilla beringei) a large silverback a few minutes before he attacked a tourist, Virunga Mountains, Rwanda

Cyril Ruoso / Mandrill (Mandrillus sphinx) adult male vocalizing showing huge canine teeth, Gabon

Michael Pitts / Stonefish (Synanceia verrucosa) close up portrait, Red Sea.

Loic Poidevin / Short eared owl (Asio flammeus) in grass, Breton marsh, West France, January

Hermann Brehm / Jaguar (Panthera onca) female snarling, Pantanal, Brazil

Anup Shah / Western chimpanzee (Pan troglodytes verus) female 'Fana' aged 54 years protecting her grandson 'Flanle' aged 3 years from an aggressive female 'Yo' aged 49 years who is angry at a juvenile female 'J...

Mike Wilkes / Short-eared Owl (Asio flammeus) perched on rock. Genovesa, Galapagos Islands.

Mark Taylor / Yellow Labrador Retriever puppy, 8 weeks, with Lionhead cross rabbit, Tedson. NON DISPONIBLE POUR UNE UTILISATION DANS UN LIVRE JUSQU'EN 2025. MERCI DE NOUS CONTACTER POUR TOUT AUTRE UTILISATION AFIN...

Charlie Summers / Angry clouds over the Central Kalahari just prior to a torrential rain and lightning storm hit, Botswana

Michael D. Kern / Diamondback terrapin, (Malaclemys terrapin) captive from USA

Martin Willis / Blue-winged Kookaburra (Dacelo leachii) male in nest with hatchlings begging, Townsville, Queensland, Australia

Anup Shah / Lioness snarling (Panthera leo) Masai Mara National Reserve, Kenya

Anup Shah / Lioness snarling (Panthera leo) Masai Mara National Reserve, Kenya

Anup Shah / Lioness snarling behind male (Panthera leo) Masai Mara National Reserve, Kenya

Lassi Rautiainen / Bean goose (Anser fabalis) walking over snow, crouched low, hissing, aggresion, Finland, May

Daniel Heuclin / Portrait of a Desert Tortoise (Gopherus agassizii). Controlled conditions. Foothills of Catalina mountains, Arizona, October.

Suzi Eszterhas / Infant Chimpanzees (Pan troglodytes) playing in tree. Tropical forest, Western Uganda

Stephen Belcher / Mountain Gorilla (Gorilla gorilla beringei), Virunga Mountains, Volcanoes National Park, Rwanda

Martin van Lokven / Leopard (Panthera pardus) female, Serengeti National Park, Tanzania

Konrad Wothe / Siberian Tiger (Panthera tigris altaica), captive individual in snow snarling, endangered species native to Asia

Thomas Marent / Mountain Gorilla (Gorilla gorilla beringei) silverback displaying, Volcanoes National Park, Rwanda

Thomas Marent / Mountain Gorilla (Gorilla gorilla beringei) silverback displaying by beating chest and calling, Volcanoes National Park, Rwanda

Thomas Marent / Mountain Gorilla (Gorilla gorilla beringei) silverback displaying by beating chest and calling, Volcanoes National Park, Rwanda

Thomas Marent / Mountain Gorilla (Gorilla gorilla beringei) silverback displaying, Volcanoes National Park, Rwanda

Tim Fitzharris / Gray Wolf (Canis lupus) snarling, North America

Konrad Wothe / Northern Elephant Seal (Mirounga angustirostris) juvenile males play fighting, North America

Suzi Eszterhas / Cheetah (Acinonyx jubatus) hissing, Cheetah Conservation Fund, Namibia

Konrad Wothe / Grizzly Bear (Ursus arctos horribilis) calling, Colorado

Michael D. Kern / King Cobra {Ophiophagus hannah} portrait tasting with tongue, captive, South eastern Asia snake

Konrad Wothe / Domestic Cat (Felis catus) in deep snow, Germany

Anup Shah / 11 yr aggressive male Sumatran orang utan (Pongo abelii) Gunung Leuser NP, Sumatra, Indonesia

Jeff Vanuga / Male African lion {Panthera leo} snarling, Samburu National Reserve, Kenya, Africa.

Jorma Luhta / Brambling male aggressive posture {Fringilla montifringilla} Finland

Matthias Breiter / Grizzly Bear (Ursus arctos horribilis) mother defending her offspring from another bear, Katmai National Park, Alaska

Shin Yoshino / Hippopotamus (Hippopotamus amphibius) pair fighting, Masai Mara, Kenya

Winfried Wisniewski / Cheetah (Acinonyx jubatus) snarling, Africa

Richard Du Toit / Leopard (Panthera pardus) adult female snarling, summer, Moremi Wildlife Reserve, Botswana

Hiroya Minakuchi / Southern Elephant Seal (Mirounga leonina) male roaring, Falkland Islands

Claus Meyer / Humboldt's Woolly Monkey (Lagothrix lagotricha) calling from tree, Amazon ecosystem, Brazil

Konrad Wothe / Siberian Tiger (Panthera tigris altaica) growling in snow, Siberian Tiger Park, Harbin, China

Tim Fitzharris / Grizzly Bear (Ursus arctos horribilis) calling, North America

Norbert Wu / Blue-ringed Octopus (Hapalochlaena sp) deadly venom can paralyze a human's respiratory system in eight minutes, pulses blue rings when angry, Australia

Gerry Ellis / Northern Elephant Seal (Mirounga angustirostris) males fighting, Ano Nuevo, California

Tim Fitzharris / Grizzly Bear (Ursus arctos horribilis) calling, North America

Mitsuaki Iwago / Domestic Cat (Felis catus) kitten hissing, Japan

Jim Brandenburg / Mute Swan (Cygnus olor) males fighting, Sweden

ZSSD / Malayan Tiger (Panthera tigris jacksoni) snarling, native to Malaysia

Geoff Simpson / Noctule bat {Nyctalus noctula} Derbyshire UK

Bengt Lundberg / Field vole showing teeth {Microtus agrestis} Sweden

Anup Shah / Annoyed male tiger snarling. (Panthera tigris) Ranthambhore NP India

Suzi Eszterhas / Tasmanian Devil (Sarcophilus harrisii) in defensive posture, Trowunna Wildlife Sanctuary, Tasmania, Australia

Suzi Eszterhas / Tasmanian Devil (Sarcophilus harrisii) in defensive posture, Trowunna Wildlife Sanctuary, Tasmania, Australia

Heike Odermatt / Southern Elephant Seal (Mirounga leonina) pup calling during defensive display, Sea Lion Island, Falkland Islands

Sumio Harada / Coyote (Canis latrans) snarling in snow, Yellowstone National Park, Montana

Sergey Gorshkov / Arctic Fox (Alopex lagopus) carrying an egg away from an angry adult Snow Goose (Chen caerulescens), Wrangel Island, Russia

Paul D Stewart / Copperplate engraving of mythical hydra with hand colouring by J. Chapman 1806 after engraving by Seba in his 'Treasury of Natural History' (1734). In 1735 a young Linnaeus visited Hamburg. While ther...

Winfried Wisniewski / Cheetah (Acinonyx jubatus) in aggressive posture, Africa

Richard Du Toit / Cape Weaver (Ploceus capensis) male, Garden Route National Park, South Africa