Theo Bosboom / Sand patterns created by receding waves, Quiberon, Brittany, France. August 2017
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Alex Mustard / Orange painted frogfish (Antennarius pictus) fishing. Here the frogfish is illuminated by a snooted strobe and the background sand lit with a red LED torch. Bitung, North Sulawesi, Indonesia. Lembeh S...
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Alex Mustard / Whitemargin stargazer (Uranoscopus sulphureus) sets an ambush in the sand at night. Bitung, North Sulawesi, Indonesia. Lembeh Strait, Molucca Sea.
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Nick Upton / Sand-loving / Square-headed wasp (Tachysphex costae) feeding on Sea holly flowers (Eryngium maritimum) on a beach, Sardinia, Italy, June.
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Milan Radisics / Aerial shot of the Hafursa glacial river running over black sand, southern Iceland, August 2018.
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Milan Radisics / Aerial shot of a confluence of three tributaries flowing over black sand at the mouth of the Affall River, Landeyjar area, Southern Iceland, August 2018.
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John Chan / Killdeer (Charadrius vociferus) stamping its feet to create vibrations in the sand as it forages for insects, Southern California, USA, August.
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David Woodfall / Cheviot sheep grazing eroding machair at front of sand dunes, North Uist, Scotland, UK, June.
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David Woodfall / Little tern (Sterna albifrons ) feeding sand eel (Hyperoplus spp) to young chick. Gronant Dunes, Denbighshire, Wales, UK, June.
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David Woodfall / Sequence of strips of cultivated Machair with early shoots of Black oats (Avena strigosa) pushing through the sand. North Uist, Scotland, UK, June.
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Daniel Heuclin / Horned viper (Vipera ammodytes) in natural setting, Turkey.
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Terry Whittaker / Red foxes (Vulpes vulpes) fighting in sand dune habitat, Holland, Netherlands. November.