Sergey Gorshkov / Lionesses (Panthera leo) trying to bring down African buffalo (Syncerus caffer) Londolozi Private Game Reserve, Sabi Sand Game Reserve, South Africa. Sequence 5 of 5

Laurent Geslin / Pride of Lions walking along road followed by tourist vehicles, Kruger National Park

Denis-Huot,Denis-Huot / Lion (Panthera leo) pride lying in long grass watching elephants (Loxodonta africana), Masai-Mara Game Reserve, Kenya. Vulnerable species.

Anup Shah / Defassa waterbuck (Kobus ellipsiprymnus defassa) male standing in a lake surrounded by a pride of African lions (Panthera leo), Masai Mara National Reserve, Kenya, September sequence 8/11

Anup Shah / Defassa waterbuck (Kobus ellipsiprymnus defassa) male standing in a lake surrounded by a pride of African lions (Panthera leo), Masai Mara National Reserve, Kenya, September sequence 1/11

Anup Shah / Defassa waterbuck (Kobus ellipsiprymnus defassa) male standing in a lake surrounded by a pride of African lions (Panthera leo), Masai Mara National Reserve, Kenya, September sequence 7/11

Anup Shah / Eastern White bearded Wildebeest (Connochaetes taurinus) being attacked by pride of African lions (Panthera leo) Masai Mara National Reserve, Kenya, September, sequence 9/12

Charlie Summers / African lion (Panthera leo) male roaring to locate the rest of his pride, Botswana, Southern Africa

Martin Dohrn / Lions (Panthera leo) approaching rhinoceros, Masai Mara, Kenya. Photographed at night using 'Starlight Camera' technology without artificial lighting.

Martin Dohrn / Pride of lions (Panthera leo) in Masai Mara, Kenya. Photographed at night using 'Starlight Camera' technology without artificial lighting.

Anup Shah / Pride of lions, adults and cubs, around tree {Panthera leo} Masai Mara Game Reserve, Kenya

Anup Shah / Pride of lions, females and cubs, walking along track in the Masai Mara, Kenya

Martin Dohrn / Pride of fifteen Lions listening for sounds of prey animals in the night. Image taken using 'Starlight Camera' technology abnd infra red light. Serengeti NP, Ngorongoro Conservation Area, Tanzania, Ea...

Martin Dohrn / Holding head low, a Gazelle watches lion pride walk by. Serengeti NP, Ngorongoro Conservation Area, Tanzania. Image taken at night using 'Starlight Camera' technology and infra red light.

Karine Aigner / Safari truck with tourists watchiung a pride of Lions (Panthera leo) as the sun rises. Ndutu area, Serengeti National Park, Tanzania

Christophe Courteau / African lions (Panthera leo) feeding at night, South Africa.

Christophe Courteau / African lions (Panthera leo) observing African elephant (Loxodonta africana) approaching them, Sabi Sand Private Game Reserve, South Africa.

Christophe Courteau / Lions (Panthera leo) observing an African elephant (Loxodonta africana) approaching them, Sabi Sand Private Game Reserve, South Africa.

Anup Shah / Lions (Panthera leo) lionesses playing with cubs aged 3-6 months with young male resting, Maasai Mara National Reserve, Kenya.

Anup Shah / Lions (Panthera leo) lionesses playing with cubs aged 3-6 months with young male resting, Maasai Mara National Reserve, Kenya.

Anup Shah / Lions (Panthera leo) lionesses playing with cubs aged 3-6 months with young male resting, Maasai Mara National Reserve, Kenya.

Wim van den Heever / Pride of Lions (Panthera leo) on termite mound at sunset, Okavango Delta, Botswana.

Denis-Huot / Pride of Lions (Panthera leo) feeding on Masai giraffe (Giraffa camelopardalis tippelskirchi) kill, Masai-Mara game reserve, Kenya.

Ammonite / Group of African lions (Panthera leo) feeding on kill, footage taken at night using thermal camera technology, Masai Mara, Kenya.

Ammonite / Pride of African lions (Panthera leo) feeding on Warthog (Phacochoerus aethiopicus) kill, Moremi Game Reserve, Botswana.

Ammonite / Group of juvenile African lions (Panthera leo) playing with rib bones from kill, footage taken at night using thermal camera technology, Moremi Game Reserve, Botswana.

Will Burrard-Lucas / Small pride of Lions (Panthera leo) on starry night. South Luangwa National Park, Zambia.

Wim van den Heever / Pride of African lions (Panthera leo) at sunrise, Okavango Delta, Botswana.

John Downer / Lion (Panthera Leo) sitting in front of 'Bouldercam' remote camera, Masai Mara, Kenya, Africa. Taken on location for 'Pride' tv series.

Anup Shah / Cape buffalo (Syncerus caffer) chasing a pride of lions (Panthera leo). Masai Mara National Reserve, Kenya, August

Anup Shah / Cape buffalo herd (Syncerus caffer) advancing on a pride of lions (Panthera leo). Masai Mara National Reserve, Kenya, August

Anup Shah / Cape buffalo herd (Syncerus caffer) advancing on a pride of lions (Panthera leo). Masai Mara National Reserve, Kenya, August

Anup Shah / Cape buffalo herd (Syncerus caffer) advancing on a pride of lions (Panthera leo). Masai Mara National Reserve, Kenya, August

Denis-Huot / Lioness (Panthera leo) nuzzling up to another female of the pride, Masai-Mara Game Reserve, Kenya. Vulnerable species.

Mark MacEwen / Marsh pride lion cubs of the Marsh pride (Panthera leo) on a moonless night, Masai Mara, Kenya, taken with infra red camera, September

Christophe Courteau / Pride of African lions (Panthera leo) with a kill at night time, watched by Brown hyaenas (Hyaena brunnea). The Dominant Lioness is defending the clan and faces the Hyaenas. Botswana

Kristel Richard / Pride of African lions sunbathing (Panthera leo) in the early morning, Mombo, Moremi Game Reserve, Chief Island, Okavango Delta, Botswana.

Anup Shah / Defassa waterbuck (Kobus ellipsiprymnus defassa) male standing in a lake surrounded by a pride of African lions (Panthera leo), Masai Mara National Reserve, Kenya, September sequence 3/11

Anup Shah / Defassa waterbuck (Kobus ellipsiprymnus defassa) male standing in a lake surrounded by a pride of African lions (Panthera leo), Masai Mara National Reserve, Kenya, September sequence 2/11

Anup Shah / Defassa waterbuck (Kobus ellipsiprymnus defassa) male standing in a lake surrounded by a pride of African lions (Panthera leo), Masai Mara National Reserve, Kenya, September sequence 4/11

Anup Shah / Defassa waterbuck (Kobus ellipsiprymnus defassa) male standing in a lake surrounded by a pride of African lions (Panthera leo), Masai Mara National Reserve, Kenya, September sequence 6/11

Anup Shah / Defassa waterbuck (Kobus ellipsiprymnus defassa) male standing in a lake surrounded by a pride of African lions (Panthera leo), Masai Mara National Reserve, Kenya, September sequence 5/11

Anup Shah / Eastern White bearded Wildebeest (Connochaetes taurinus) in stand-off with pride of African lions (Panthera leo) Masai Mara National Reserve, Kenya, September, sequence 6/12

Anup Shah / Eastern White bearded Wildebeest (Connochaetes taurinus) being attacked by pride of African lions (Panthera leo) Masai Mara National Reserve, Kenya, September, sequence 8/12

Anup Shah / Eastern White bearded Wildebeest (Connochaetes taurinus) being attacked by pride of African lions (Panthera leo) Masai Mara National Reserve, Kenya, September, sequence 10/12

Anup Shah / Eastern White bearded Wildebeest (Connochaetes taurinus) being attacked by pride of African lions (Panthera leo) Masai Mara National Reserve, Kenya, September, sequence 7/12

Anup Shah / Eastern White bearded Wildebeest (Connochaetes taurinus) being attacked by pride of African lions (Panthera leo) Masai Mara National Reserve, Kenya, September, sequence 5/12

Anup Shah / Eastern White bearded Wildebeest (Connochaetes taurinus) being attacked by pride of African lions (Panthera leo) Masai Mara National Reserve, Kenya, September, sequence 2/12

Anup Shah / Eastern White bearded Wildebeest (Connochaetes taurinus) in stand-off with pride of African lions (Panthera leo) Masai Mara National Reserve, Kenya, September, sequence 4/12

Anup Shah / Eastern White bearded Wildebeest (Connochaetes taurinus) in stand-off with pride of African lions (Panthera leo), Masai Mara National Reserve, Kenya, September, sequence 1/12

Andy Rouse / Two African lionesses (Panthera leo) lioness walking with group of cubs, Masai Mara National Reserve, Kenya

Sergey Gorshkov / African lions (Panthera leo) close up and low angle shot of pride feeding on carcass, Okavango Delta, Botswana

Sergey Gorshkov / African lion (Panthera leo) close up and low angle shot of male snatching away part of carcass from rest of pride, Okavango Delta, Botswana

Sergey Gorshkov / African lion (Panthera leo) close up view of pride feeding, lioness looking directly into lens, Okavango Delta, Botswana

Ben Osborne / African lions (Panthera leo) feed on carcass of an Elephant (Loxodonta africana) that they killed the previous night, Savuti, Northern Botswana. Taken on location for BBC Planet Earth series, 2005

Ben Osborne / African lions (Panthera leo) feed on carcass of an Elephant (Loxodonta africana) that they killed the previous night. The lion on the right is threatening new arrivals off to the left of the picture....

Ben Osborne / African lion (Panthera leo) pride of lions eats an Eland carcass under shrub for shade, Botswana, taken on location for BBC Planet Earth series, 2005

Ben Osborne / A pride of African lions (Panthera leo) feed on the carcass of an African elephant (Loxodonta africana)they hunted and killed the previous night and have returned to finish eating the meat off the car...

Andy Rouse / African lion (Panthera leo) male lion of the Marsh pride, Masai Mara GR, Kenya, February

Anup Shah / Lion pride (Panthera leo) testing a herd of Cape Buffalo (Syncerus cafer). Masai Mara National Reserve, Kenya, August 2009

Anup Shah / African lion (Panthera leo) pride resting in shade of a tree, Masai Mara National Reserve, Kenya. April.

Anup Shah / African elephant (Loxodonta africana) approaching a pride of lions (Panthera leo). Masai Mara National Reserve, Kenya. March.

Anup Shah / African elephant (Loxodonta africana) approaching a pride of lions (Panthera leo) Masai Mara National Reserve, Kenya. March.

Anup Shah / African elephant (Loxodonta africana) in stand-off with a pride of lions (Panthera leo). Masai Mara National Reserve, Kenya. March

Anup Shah / Lion pride (Panthera leo) resting in shade of a tree, watching an African elephant pass by (Loxodonta africana) Masai Mara National Reserve, Kenya. March.

Warwick Sloss / African lionesses {Panthera leo} licking each other as part of the social bonding within the pride, Katavi NP, Tanzania.

Edwin Giesbers / Pride of African lions (Panthera leo) walking in a line, Serengeti NP, Tanzania

Edwin Giesbers / Pride of African lions (Panthera leo) drinking, Serengeti NP, Tanzania

Edwin Giesbers / Pride of African lions (Panthera leo) walking along a track, Serengeti NP, Tanzania

Edwin Giesbers / Pride of African lions (Panthera leo) lying by water with some drinking, Serengeti NP, Tanzania

Edwin Giesbers / Pride of African lions (Panthera leo) drinking, Serengeti NP, Tanzania

Edwin Giesbers / Pride of African lions (Panthera leo) drinking, Tanzania

Michele Westmorland / African lionesses (Panthera leo) of the Tsaro Pride wading through water, Duba Plains, Okavango Delta, Botswana

Michele Westmorland / African lionesses (Panthera leo) of the Tsaro Pride wding through water, Duba Plains, Okavango Delta, Botswana

Michele Westmorland / Two African lionesses (Panthera leo) of the Tsaro Pride, resting on termite mound, Duba Plains, Okavango Delta, Botswana

Michele Westmorland / African lionesses (Panthera leo) of the Tsaro Pride resting on termite mound (note Cape buffalo horns from prey) Duba Plains, Okavango Delta, Botswana

Anup Shah / African lion {Panthera leo} pride resting on top of Kopje, Serengeti NP, Tanzania

Pete Oxford / Elephant herd (Loxodonta africana) approaching a waterhole where a pride of lions (Panthera leo) are drinking. Makalolo Plains, Hwange National Park, Zimbabwe, Southern Africa

Pete Oxford / Elephant herd (Loxodonta africana) approaching a waterhole where a pride of lions (Panthera leo) are drinking. Makalolo Plains, Hwange National Park, Zimbabwe, Southern Africa

Pete Oxford / Tourists watching a pride of Lions {Panthera leo} feeding on a Buffalo kill. Duba Plains, Okavango Delta, Botswana, Southern Africa 2006

Richard Du Toit / Aerial view of African lion pride (Panthera leo) hunting Buffalo (Syncerus caffer) Okavango Delta, Botswana

Anup Shah / Pack of African lions {Panthera leo} approaching a cornered African buffalo {Syncerus caffer} Masai Mara GR, Kenya

Anup Shah / Pride of lions walking - lionesses with cubs {Panthera leo} Masai Mara, Kenya

Pete Oxford / Pride of Lions {Panthera leo} feeding on Elephant carcass. Okavango, Botswana

Anup Shah / Pride of African lions {Panthera leo} resting on savanna under stormy sky, Serengeti National Park, Tanzania

Sharon Heald / Pride of African lions {Panthera leo} drinking. Namibia

Pete Oxford / Pride of lions {Panthera leo} Okavango delta, Botswana

Peter Blackwell / Head portrait of lioness in pride. Masai Mara Kenya

Carine Schrurs / Lion pride resting in shade of tree {Panthera leo} Samburu NR, Kenya

John Downer / Pride of Lions asleep in shade of tree {Panthera leo} Masai Mara, Kenya

Peter Blackwell / Pride of female Lions (Panthera leo) resting in grassland with approaching storm, Masai Mara GR, Kenya

Suzi Eszterhas / Three Lions (Panthera leo) female, walking over grassland with cubs, aged 6-12 weeks, Sabi Sands Game Reserve, South Africa.

Denis-Huot / Pride of Lions (Panthera leo) walking through tall grass during storm. Masai-Mara National Reserve, Kenya.

Charlie Hamilton-James / Three Maasai herders butchering a cow killed by the Talek pride of Lions (Panthera leo) , Masai Mara, Kenya, Africa. April, 2015.

Charlie Hamilton-James / Maasai cattle herders butchering a cow killed by the Talek pride of Lions (Panthera leo) , Masai Mara , Kenya, Africa. April, 2015.