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01721268

Ugo Mellone / Landscape of Guadix geological depression within Los Colorados desert habitat. Guadix depression, Andalusia, Spain. October.

01721267

Ugo Mellone / Iberian ibex (Capra pyrenaica), pair with male top right, during rutting season. Sierra Nevada National Park, Andalusia, Spain. October.

Landscape of Guadix geological depression within Los Colorados desert habitat, with Sierra Nevada National Park in background. Guadix depression, Andalusia, Spain. February.

01721266

Ugo Mellone / Landscape of Guadix geological depression within Los Colorados desert habitat, with Sierra Nevada National Park in background. Guadix depression, Andalusia, Spain. February.

Iberian ibex (Capra pyrenaica), adult male, walking along vertical cliff. Guadix depression, Spanish badlands, Andalusia, Spain. December.

01721265

Ugo Mellone / Iberian ibex (Capra pyrenaica), adult male, walking along vertical cliff. Guadix depression, Spanish badlands, Andalusia, Spain. December.

01721264

Ugo Mellone / Iberian ibex (Capra pyrenaica), adult male, at top of cliff. Guadix depression, Spanish badlands, Andalusia, Spain. December.

Iberian ibex (Capra pyrenaica), adult male, following female during rutting season. Guadix depression, Spanish badlands, Andalusia, Spain. December.

01721263

Ugo Mellone / Iberian ibex (Capra pyrenaica), adult male, following female during rutting season. Guadix depression, Spanish badlands, Andalusia, Spain. December.

Iberian ibex (Capra pyrenaica), adult male, behind female atop cliff during rutting season. Guadix depression, Spanish badlands, Andalusia, Spain. December.

01721262

Ugo Mellone / Iberian ibex (Capra pyrenaica), adult male, behind female atop cliff during rutting season. Guadix depression, Spanish badlands, Andalusia, Spain. December.

Iberian ibex (Capra pyrenaica), adult male, jumping between cliff faces. Guadix depression, Spanish badlands, Andalusia, Spain. December.

01721261

Ugo Mellone / Iberian ibex (Capra pyrenaica), adult male, jumping between cliff faces. Guadix depression, Spanish badlands, Andalusia, Spain. December.

01721260

Ugo Mellone / Two Iberian ibex (Capra pyrenaica) yearlings playfighting. Guadix depression, Spanish badlands, Andalusia, Spain. December.

01721259

Ugo Mellone / Two Iberian ibex (Capra pyrenaica) running down snow covered slope. Sierra Nevada National Park, Andalusia, Spain. November.

Iberian ibex (Capra pyrenaica), adult male, exhibiting flehmen response towards female during rutting season. Sierra Nevada National Park, Andalusia, Spain. November.

01721258

Ugo Mellone / Iberian ibex (Capra pyrenaica), adult male, exhibiting flehmen response towards female during rutting season. Sierra Nevada National Park, Andalusia, Spain. November.

01721257

Ugo Mellone / Iberian ibex (Capra pyrenaica) herd, all male, walking along coastal cliffs. Mediterranean coast, Andalusia, Spain. December.

01721256

Ugo Mellone / Iberian ibex (Capra pyrenaica) herd, all male, walking along coastal cliffs. Mediterranean coast, Andalusia, Spain. December.

01721255

Ugo Mellone / Iberian ibex (Capra pyrenaica), female, feeding from bush growing on cliff side. Sierra Nevada National Park, Andalusia, Spain. May.

01721254

Ugo Mellone / Iberian ibex (Capra pyrenaica), adult male, portrait, at 2900m altitude. Sierra Nevada National Park, Andalusia, Spain. July.

01721253

Ugo Mellone / Iberian ibex (Capra pyrenaica), adult male, portrait, at 2900m altitude. Sierra Nevada National Park, Andalusia, Spain. July.

Iberian ibex (Capra pyrenaica), adult male, shaking off dust and scratching, behind cloud of gnats, at 2900m altitude. Sierra Nevada National Park, Andalusia, Spain. July.

01721252

Ugo Mellone / Iberian ibex (Capra pyrenaica), adult male, shaking off dust and scratching, behind cloud of gnats, at 2900m altitude. Sierra Nevada National Park, Andalusia, Spain. July.

Iberian ibex (Capra pyrenaica) kid, aged two months, resting on his mother's neck, at 2900 m altitude. Sierra Nevada National Park, Andalusia, Spain. July.

01721251

Ugo Mellone / Iberian ibex (Capra pyrenaica) kid, aged two months, resting on his mother's neck, at 2900 m altitude. Sierra Nevada National Park, Andalusia, Spain. July.

Iberian ibex (Capra pyrenaica), female, feeding while kid, aged between one and two months, pesters. Sierra Nevada National Park, Andalusia, Spain. May.

01721250

Ugo Mellone / Iberian ibex (Capra pyrenaica), female, feeding while kid, aged between one and two months, pesters. Sierra Nevada National Park, Andalusia, Spain. May.

01721249

Ugo Mellone / Iberian ibex (Capra pyrenaica), adult males, running down slope, at 2900m altitude. Sierra Nevada National Park, Andalusia, Spain. July.

Iberian ibex (Capra pyrenaica) nursery, composed of females and kids, feeding along cliff face. Sierra Nevada National Park, Andalusia, Spain. May.

01721248

Ugo Mellone / Iberian ibex (Capra pyrenaica) nursery, composed of females and kids, feeding along cliff face. Sierra Nevada National Park, Andalusia, Spain. May.

01721247

Ugo Mellone / Iberian ibex (Capra pyrenaica) kid playfighting, at 2900m altitude. Sierra Nevada National Park, Andalusia, Spain. July.

01721246

Ugo Mellone / Two Iberian ibex (Capra pyrenaica), adult males, fighting. Sierra Nevada National Park, Andalusia, Spain. July.

01721245

Ugo Mellone / Two Iberian ibex (Capra pyrenaica), adult males, fighting. Sierra Nevada National Park, Andalusia, Spain. July.

01721244

Ugo Mellone / Iberian ibex (Capra pyrenaica), adult male, feeding. Sierra Nevada National Park, Andalusia, Spain. July.

01721243

Ugo Mellone / Photographer's campsite during fieldwork on Iberian ibex. Sierra Nevada National Park, Andalusia, Spain. July.

01721242

Ugo Mellone / Photographer and his campsite during fieldwork on Iberian ibex in Siete Lagunas. Sierra Nevada National Park, Andalusia, Spain. July.

Iberian ibex (Capra pyrenaica), adult males, grazing along stream formed from glacial melt, at 2900m altitude. Sierra Nevada National Park, Andalusia, Spain. July.

01721241

Ugo Mellone / Iberian ibex (Capra pyrenaica), adult males, grazing along stream formed from glacial melt, at 2900m altitude. Sierra Nevada National Park, Andalusia, Spain. July.

01721240

Ugo Mellone / Iberian ibex (Capra pyrenaica), adult female, hooves and legs close up. Sierra Nevada National Park, Andalusia, Spain. June.

Landscape of Sierra Nevada's highest snow capped peaks with lenticular clouds over them. Sierra Nevada National Park, Andalusia, Spain. March.

01721239

Ugo Mellone / Landscape of Sierra Nevada's highest snow capped peaks with lenticular clouds over them. Sierra Nevada National Park, Andalusia, Spain. March.

Iberian ibex (Capra pyrenaica), adult male, exhibiting flehmen response during rutting season. Sierra Nevada National Park, Andalusia, Spain. November.

01721238

Ugo Mellone / Iberian ibex (Capra pyrenaica), adult male, exhibiting flehmen response during rutting season. Sierra Nevada National Park, Andalusia, Spain. November.

01721237

Ugo Mellone / Two Iberian ibex (Capra pyrenaica), adult males, fighting. Sierra Nevada National Park, Andalusia, Spain. June.

01721236

Ugo Mellone / Iberian ibex (Capra pyrenaica), adult male, licking salt, at 2900m altitude. Sierra Nevada National Park, Andalusia, Spain. July.

01721235

Ugo Mellone / Iberian ibex (Capra pyrenaica), adult male, portrait. Sierra Nevada National Park, Andalusia, Spain. June.

01721234

Ugo Mellone / Iberian ibex (Capra pyrenaica), adult male, shadow. Trevelez valley, Sierra Nevada National Park, Andalusia, Spain. June.

01721233

Ugo Mellone / Iberian ibex (Capra pyrenaica), adult male, horns close up. Sierra Nevada National Park, Andalusia, Spain. June.

Landscape of Spanish badlands, with Iberian ibex (Capra pyrenaica) grazing in foreground. Sierra Nevada National Park, Andalusia, Spain. June.

01721232

Ugo Mellone / Landscape of Spanish badlands, with Iberian ibex (Capra pyrenaica) grazing in foreground. Sierra Nevada National Park, Andalusia, Spain. June.

01721231

Ugo Mellone / Iberian ibex (Capra pyrenaica), adult male, walking along mountain ridge. Sierra Nevada National Park, Andalusia, Spain. June.

Photographer's campsite equipped with solar panels, during fieldwork on Iberian ibex. Sierra Nevada National Park, Andalusia, Spain. June.

01721230

Ugo Mellone / Photographer's campsite equipped with solar panels, during fieldwork on Iberian ibex. Sierra Nevada National Park, Andalusia, Spain. June.

Iberian ibex (Capra pyrenaica) mixed herd crossing mountain pass after snowfall. Sierra Nevada National Park, Andalusia, Spain. February.

01721229

Ugo Mellone / Iberian ibex (Capra pyrenaica) mixed herd crossing mountain pass after snowfall. Sierra Nevada National Park, Andalusia, Spain. February.

01721228

Ugo Mellone / Autumn landscape of Dehesa del Camarate maple (Acer) forest. Sierra Nevada National Park, Andalusia, Spain. October.

Iberian ibex (Capra pyrenaica), adult males, grazing beside glacial lake, at 2900m altitude. Sierra Nevada National Park, Andalusia, Spain. June.

01721227

Ugo Mellone / Iberian ibex (Capra pyrenaica), adult males, grazing beside glacial lake, at 2900m altitude. Sierra Nevada National Park, Andalusia, Spain. June.

01721226

Ugo Mellone / Iberian ibex (Capra pyrenaica) kid peering over rocks. Sierra Nevada National Park, Andalusia, Spain. June.

01721225

Ugo Mellone / Iberian ibex (Capra pyrenaica) kid peering over rocks. Sierra Nevada National Park, Andalusia, Spain. June.

01721224

Ugo Mellone / Iberian ibex (Capra pyrenaica) kid standing on rock. Sierra Nevada National Park, Andalusia, Spain. June.

01721223

Ugo Mellone / Iberian ibex (Capra pyrenaica) kid, aged two months, at 2900m altitude. Sierra Nevada National Park, Andalusia, Spain. June.

Iberian ibex (Capra pyrenaica), adult male, exhibiting flehmen response towards two females during rutting season on coastal cliffs. Mediterranean coast, Andalusia, Spain. November.

01721222

Ugo Mellone / Iberian ibex (Capra pyrenaica), adult male, exhibiting flehmen response towards two females during rutting season on coastal cliffs. Mediterranean coast, Andalusia, Spain. November.

01721221

Ugo Mellone / Iberian ibex (Capra pyrenaica), adult male, walking along cliff edge. Guadix depression, Spanish badlands, Andalusia, Spain. February.

Iberian ibex (Capra pyrenaica) twins, aged between one and two months, suckling from mother. Sierra Nevada National Park, Andalusia, Spain. May.

01721220

Ugo Mellone / Iberian ibex (Capra pyrenaica) twins, aged between one and two months, suckling from mother. Sierra Nevada National Park, Andalusia, Spain. May.

01721219

Ugo Mellone / Two Iberian ibex (Capra pyrenaica) walking along mountain ridge. Sierra Magina National Park, Andalusia, Spain. October.

01721218

Ugo Mellone / Iberian ibex (Capra pyrenaica), adult male, calling, at 2900m altitude. Sierra Nevada National Park, Andalusia, Spain. June.

01721217

Ugo Mellone / Iberian ibex (Capra pyrenaica), adult male, resting and grooming, at 2900 m altitude. Sierra Nevada National Park, Andalusia, Spain. June.

01721216

Ugo Mellone / Iberian ibex (Capra pyrenaica) kids, aged between one and two months, playfighting. Sierra Nevada National Park, Andalusia, Spain. May.

Iberian ibex (Capra pyrenaica), adult male, walking along snow covered mountain ridge. Sierra Nevada National Park, Andalusia, Spain. February.

01721215

Ugo Mellone / Iberian ibex (Capra pyrenaica), adult male, walking along snow covered mountain ridge. Sierra Nevada National Park, Andalusia, Spain. February.

Iberian ibex (Capra pyrenaica) kids, aged between one and two months, climbing over rocks. Sierra Nevada National Park, Andalusia, Spain. May.

01721214

Ugo Mellone / Iberian ibex (Capra pyrenaica) kids, aged between one and two months, climbing over rocks. Sierra Nevada National Park, Andalusia, Spain. May.

Iberian ibex (Capra pyrenaica), adult male with yearlings, climbing over cliff, at 2900m altitude. Sierra Nevada National Park, Andalusia, Spain. June.

01721213

Ugo Mellone / Iberian ibex (Capra pyrenaica), adult male with yearlings, climbing over cliff, at 2900m altitude. Sierra Nevada National Park, Andalusia, Spain. June.

Iberian ibex (Capra pyrenaica), adult female, grazing with kid, aged between one and two months. Sierra Nevada National Park, Andalusia, Spain. May.

01721212

Ugo Mellone / Iberian ibex (Capra pyrenaica), adult female, grazing with kid, aged between one and two months. Sierra Nevada National Park, Andalusia, Spain. May.

Iberian ibex (Capra pyrenaica) yearling making its way down snow covered slope. Sierra Nevada National Park, Andalusia, Spain. February.

01721211

Ugo Mellone / Iberian ibex (Capra pyrenaica) yearling making its way down snow covered slope. Sierra Nevada National Park, Andalusia, Spain. February.

Iberian ibex (Capra pyrenaica), adult male, in snow covered landscape after snowfall. Sierra Nevada National Park, Andalusia, Spain. February.

01721210

Ugo Mellone / Iberian ibex (Capra pyrenaica), adult male, in snow covered landscape after snowfall. Sierra Nevada National Park, Andalusia, Spain. February.

Iberian ibex (Capra pyrenaica), adult female with yearling, in snow covered landscape after snowfall. Sierra Nevada National Park, Andalusia, Spain. February.

01721209

Ugo Mellone / Iberian ibex (Capra pyrenaica), adult female with yearling, in snow covered landscape after snowfall. Sierra Nevada National Park, Andalusia, Spain. February.

01721208

Ugo Mellone / Iberian ibex (Capra pyrenaica), adult female with yearling, in snow covered landscape after snowfall. Sierra Nevada National Park, Andalusia, Spain. February.

Landscape of mountain range at sunset, with Trevenque peak in centre, during winter. Sierra Nevada National Park, Andalusia, Spain. February.

01721207

Ugo Mellone / Landscape of mountain range at sunset, with Trevenque peak in centre, during winter. Sierra Nevada National Park, Andalusia, Spain. February.

01721206

Ugo Mellone / Iberian ibex (Capra pyrenaica), adult male, beside glacial lake, at 2900m altitude. Sierra Nevada National Park, Andalusia, Spain. June.

01721205

Ugo Mellone / Landscape of snow covered mountain range. Sierra Nevada National Park, Andalusia, Spain. February.

Two Iberian ibex (Capra pyrenaica), adult males, grazing beside glacial lake, at 2900m altitude. Sierra Nevada National Park, Andalusia, Spain. June.

01721204

Ugo Mellone / Two Iberian ibex (Capra pyrenaica), adult males, grazing beside glacial lake, at 2900m altitude. Sierra Nevada National Park, Andalusia, Spain. June.

Iberian ibex (Capra pyrenaica), two adult females with kid aged between one and two months, climbing over rocks. Sierra Nevada National Park, Andalusia, Spain. May.

01721203

Ugo Mellone / Iberian ibex (Capra pyrenaica), two adult females with kid aged between one and two months, climbing over rocks. Sierra Nevada National Park, Andalusia, Spain. May.

01721202

Ugo Mellone / Iberian ibex (Capra pyrenaica) tracks in snow. Sierra Nevada National Park, Andalusia, Spain. February.

01721201

Ugo Mellone / Borreguiles, typical wet meadows, beside Mosca glacial lake, at 2900m altitude. Sierra Nevada National Park, Andalusia, Spain. June.

Three Iberian ibex (Capra pyrenaica), adult males, standing on rocks, at 2900m altitude. Sierra Nevada National Park, Andalusia, Spain. July.

01721200

Ugo Mellone / Three Iberian ibex (Capra pyrenaica), adult males, standing on rocks, at 2900m altitude. Sierra Nevada National Park, Andalusia, Spain. July.

Two Iberian ibex (Capra pyrenaica), adult males, resting beside stream, at 2900m altitude. Sierra Nevada National Park, Andalusia, Spain. July.

01721199

Ugo Mellone / Two Iberian ibex (Capra pyrenaica), adult males, resting beside stream, at 2900m altitude. Sierra Nevada National Park, Andalusia, Spain. July.

Two Iberian ibex (Capra pyrenaica), adult males, walking across high altitude rocky terrain, at 2900m altitude. Sierra Nevada National Park, Andalusia, Spain. July.

01721198

Ugo Mellone / Two Iberian ibex (Capra pyrenaica), adult males, walking across high altitude rocky terrain, at 2900m altitude. Sierra Nevada National Park, Andalusia, Spain. July.

01721197

Ugo Mellone / Last specimen of Pyrenean subspecies, bucardo, of Iberian ibex (Capra pyrenaica) housed in museum. Torla, Aragon, Spain. August.

Iberian ibex (Capra pyrenaica), adult male, crossing rocky terrain, at 2900m altitude. Sierra Nevada National Park, Andalusia, Spain. July.

01721196

Ugo Mellone / Iberian ibex (Capra pyrenaica), adult male, crossing rocky terrain, at 2900m altitude. Sierra Nevada National Park, Andalusia, Spain. July.

Iberian ibex (Capra pyrenaica), adult male, reflection in glacial lake, at 2900 m altitude. Sierra Nevada National Park, Andalusia, Spain. June.

01721195

Ugo Mellone / Iberian ibex (Capra pyrenaica), adult male, reflection in glacial lake, at 2900 m altitude. Sierra Nevada National Park, Andalusia, Spain. June.

Iberian ibex (Capra pyrenaica), adult female, resting, being pestered by kid, aged two months, begging for milk. Sierra Nevada National Park, Andalusia, Spain. July.

01721194

Ugo Mellone / Iberian ibex (Capra pyrenaica), adult female, resting, being pestered by kid, aged two months, begging for milk. Sierra Nevada National Park, Andalusia, Spain. July.

01721193

Ugo Mellone / Iberian ibex (Capra pyrenaica), adult male, portrait, at 2900 m altitude. Sierra Nevada National Park, Andalusia, Spain. July.

01721192

Ugo Mellone / Iberian ibex (Capra pyrenaica), adult male, portrait, at 2900 m altitude. Sierra Nevada National Park, Andalusia, Spain. July.

Two Iberian ibex (Capra pyrenaica), adult males, crossing rocky terrain during snowfall. Sierra Nevada National Park, Andalusia, Spain. January.

01721191

Ugo Mellone / Two Iberian ibex (Capra pyrenaica), adult males, crossing rocky terrain during snowfall. Sierra Nevada National Park, Andalusia, Spain. January.

Iberian ibex (Capra pyrenaica) herd grazing in snow covered landscape after snowfall. Sierra Nevada National Park, Andalusia, Spain. January.

01721190

Ugo Mellone / Iberian ibex (Capra pyrenaica) herd grazing in snow covered landscape after snowfall. Sierra Nevada National Park, Andalusia, Spain. January.

01721189

Ugo Mellone / Iberian ibex (Capra pyrenaica), adult female with yearling, peering out. Mediterranean coast, Andalusia, Spain. January.

Iberian ibex (Capra pyrenaica), most represented species in rock art on wall of Cueva de la Pileta, dating back 10,000 years. Andalusia, Spain. July.

01721188

Ugo Mellone / Iberian ibex (Capra pyrenaica), most represented species in rock art on wall of Cueva de la Pileta, dating back 10,000 years. Andalusia, Spain. July.

01721187

Ugo Mellone / Iberian ibex (Capra pyrenaica), most represented species in rock art on wall of Cueva de la Pileta, dating back 10,000 years. Andalusia, Spain. July.

Iberian ibex (Capra pyrenaica), female, being pestered by kid, aged two months, begging for milk. Sierra Nevada National Park, Andalusia, Spain. July.

01721186

Ugo Mellone / Iberian ibex (Capra pyrenaica), female, being pestered by kid, aged two months, begging for milk. Sierra Nevada National Park, Andalusia, Spain. July.

01721185

Ugo Mellone / Two Iberian ibex (Capra pyrenaica), adult males, fighting. Sierra Nevada National Park, Andalusia, Spain. July.

01721184

Ugo Mellone / Two Iberian ibex (Capra pyrenaica), adult males, fighting in rocky landscape. Sierra Nevada National Park, Andalusia, Spain. April.

Iberian ibex (Capra pyrenaica) mixed herd in snow covered landscape after snowfall. Sierra Nevada National Park, Andalusia, Spain. January.

01721183

Ugo Mellone / Iberian ibex (Capra pyrenaica) mixed herd in snow covered landscape after snowfall. Sierra Nevada National Park, Andalusia, Spain. January.

Two Iberian ibex (Capra pyrenaica), adult males, participating in dominance display, mounting each other. Sierra Nevada National Park, Andalusia, Spain. April.

01721182

Ugo Mellone / Two Iberian ibex (Capra pyrenaica), adult males, participating in dominance display, mounting each other. Sierra Nevada National Park, Andalusia, Spain. April.

01721181

Ugo Mellone / Two Iberian ibex (Capra pyrenaica), adult males, fighting. Sierra Nevada National Park, Andalusia, Spain. April.

01721180

Ugo Mellone / Two Iberian ibex (Capra pyrenaica), adult males, walking up from stream. Sierra Nevada National Park, Andalusia, Spain. April.

Two Iberian ibex (Capra pyrenaica) kids, aged two months, playing and jumping between rocks, at 2900m altitude. Sierra Nevada National Park, Andalusia, Spain. July.

01721179

Ugo Mellone / Two Iberian ibex (Capra pyrenaica) kids, aged two months, playing and jumping between rocks, at 2900m altitude. Sierra Nevada National Park, Andalusia, Spain. July.

01721178

Ugo Mellone / Iberian ibex (Capra pyrenaica) hooves close up. Sierra Nevada National Park, Andalusia, Spain. July.

Iberian ibex (Capra pyrenaica) kid, aged two months, making its way down cliff, at 2900m altitude. Sierra Nevada National Park, Andalusia, Spain. July.

01721177

Ugo Mellone / Iberian ibex (Capra pyrenaica) kid, aged two months, making its way down cliff, at 2900m altitude. Sierra Nevada National Park, Andalusia, Spain. July.

01721176

Ugo Mellone / Two Iberian ibex (Capra pyrenaica), adult males, fighting. Sierra Nevada National Park, Andalusia, Spain. April.

01721175

Ugo Mellone / Iberian ibex (Capra pyrenaica) at dusk. Sierra Nevada National Park, Andalusia, Spain. July.

01721174

Ugo Mellone / Iberian ibex (Capra pyrenaica), adult male, portrait between grasses. Sierra Nevada National Park, Andalusia, Spain. April.

01721173

Ugo Mellone / Iberian ibex (Capra pyrenaica) running across rocky terrain. Sierra Nevada National Park, Andalusia, Spain. July.