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Acacia tree amongst flowering Tribulus on Kalahari Dune. Kgalagadi Transfrontier Park, Southern Africa. January 2010.
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Hougaard Malan / Acacia tree amongst flowering Tribulus on Kalahari Dune. Kgalagadi Transfrontier Park, Southern Africa. January 2010.
Eland (Taurotragus oryx) males locking horns and fighting. Devil's Thorn flowers (Tribulus terrestris) cover the ground. Etosha National Park, Namibia, January.
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Tony Heald / Eland (Taurotragus oryx) males locking horns and fighting. Devil's Thorn flowers (Tribulus terrestris) cover the ground. Etosha National Park, Namibia, January.
Eland (Taurotragus oryx) male horning the ground which is covered in Devil's Thorn flowers (Tribulus terrestris). Etosha National Park, Namibia, January.
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Tony Heald / Eland (Taurotragus oryx) male horning the ground which is covered in Devil's Thorn flowers (Tribulus terrestris). Etosha National Park, Namibia, January.
Spotted Hyaena (Crocuta crocuta) rolling in Devil's Thorn flowers (Tribulus terrestris). Etosha National Park, Namibia, January.
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Tony Heald / Spotted Hyaena (Crocuta crocuta) rolling in Devil's Thorn flowers (Tribulus terrestris). Etosha National Park, Namibia, January.
Black-faced Impala (Aepyceros melamis petersi) male standing in a field of Devil's Thorn flowers (Tribulus terrestris). Etosha National Park, Namibia, January.
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Sharon Heald / Black-faced Impala (Aepyceros melamis petersi) male standing in a field of Devil's Thorn flowers (Tribulus terrestris). Etosha National Park, Namibia, January.
Zebra (Equus quagga) foal running through Devil's Thorn flowers (Tribulus terrestris). Etosha National Park, Namibia, January.
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Tony Heald / Zebra (Equus quagga) foal running through Devil's Thorn flowers (Tribulus terrestris). Etosha National Park, Namibia, January.
Devil's thorn (Tribulus zeyheri) the Kalahari desert's prominent creeper, appearing after the rains, Kgalagadi NP, South Africa
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Philippe Clement / Devil's thorn (Tribulus zeyheri) the Kalahari desert's prominent creeper, appearing after the rains, Kgalagadi NP, South Africa
Waved Albatross (Phoebastria irrorata) pair bonding in flowering tribulus field during rains, Punta Cevallos, Espanola Island, Galapagos Islands, Ecuador
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Tui De Roy / Waved Albatross (Phoebastria irrorata) pair bonding in flowering tribulus field during rains, Punta Cevallos, Espanola Island, Galapagos Islands, Ecuador
Caltrop murex (Murex Tribulus) laying their egg masses on muddy seafloor of the Lembeh Lembeh Strait, North Sulawesi, Indonesia.
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Constantinos Petrinos / Caltrop murex (Murex Tribulus) laying their egg masses on muddy seafloor of the Lembeh Lembeh Strait, North Sulawesi, Indonesia.
Yellow devil's thorn flower (Tribulus zeyheri) in the Kalahari Desert, South Africa
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Jen Guyton / Yellow devil's thorn flower (Tribulus zeyheri) in the Kalahari Desert, South Africa
Meerkat (Suricata suricatta) standing among a field of Devil's thorn flowers (Tribulus zeyheri) in the Kalahari Desert, South Africa.
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Jen Guyton / Meerkat (Suricata suricatta) standing among a field of Devil's thorn flowers (Tribulus zeyheri) in the Kalahari Desert, South Africa.
Meerkats (Suricata suricatta) two look out over a field of Devil's thorn flowers (Tribulus zeyheri) in the Kalahari Desert, South Africa.
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Jen Guyton / Meerkats (Suricata suricatta) two look out over a field of Devil's thorn flowers (Tribulus zeyheri) in the Kalahari Desert, South Africa.
Meerkats (Suricata suricatta) four looking out over a field of Devil's thorn flowers (Tribulus zeyheri) Kalahari Desert, South Africa.
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Jen Guyton / Meerkats (Suricata suricatta) four looking out over a field of Devil's thorn flowers (Tribulus zeyheri) Kalahari Desert, South Africa.
Meerkat pup (Suricata suricatta) standing among its family in a field of Devilthorn flowers (Tribulus zeyheri). Kalahari Desert, South Africa.
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Jen Guyton / Meerkat pup (Suricata suricatta) standing among its family in a field of Devilthorn flowers (Tribulus zeyheri). Kalahari Desert, South Africa.
African monarch butterfly (Danaus chrysippus) sipping nectar from a Devil's thorn flower (Tribulus zeyheri) in the Kalahari Desert, South Africa.
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Jen Guyton / African monarch butterfly (Danaus chrysippus) sipping nectar from a Devil's thorn flower (Tribulus zeyheri) in the Kalahari Desert, South Africa.
Egyptian goose (Alopochen aegyptiaca) pair amongst Devil's-thorn yellow flowers  (Tribulus terrestris) Etosha National Park, Namibia, March.
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Tony Heald / Egyptian goose (Alopochen aegyptiaca) pair amongst Devil's-thorn yellow flowers (Tribulus terrestris) Etosha National Park, Namibia, March.
White-backed vulture (Gyps africanus) sunbathing amongst Devil's-thorn yellow flowers (Tribulus terrestris), Etosha National Park, Namibia, March.
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Tony Heald / White-backed vulture (Gyps africanus) sunbathing amongst Devil's-thorn yellow flowers (Tribulus terrestris), Etosha National Park, Namibia, March.
Klein Okevi waterhole covered by Devil's-thorn yellow flowers  (Tribulus terrestris) Etosha National Park, Namibia, March 2016
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Tony Heald / Klein Okevi waterhole covered by Devil's-thorn yellow flowers (Tribulus terrestris) Etosha National Park, Namibia, March 2016
Black-faced impala (Aepyceros melampus petersi) young male Lying in Devil's-thorn yellow flowers  (Tribulus terrestris) Etosha National Park  Namibia, March.
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Tony Heald / Black-faced impala (Aepyceros melampus petersi) young male Lying in Devil's-thorn yellow flowers (Tribulus terrestris) Etosha National Park Namibia, March.
Black-faced impala (Aepyceros melampus petersi) female leaping above Devil's-thorn yellow flowers  (Tribulus terrestris) Etosha National Park, Namibia.
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Tony Heald / Black-faced impala (Aepyceros melampus petersi) female leaping above Devil's-thorn yellow flowers (Tribulus terrestris) Etosha National Park, Namibia.
Lion (Panthera leo) male, walking amongst Devil's-thorn yellow flowers (Tribulus terrestris) Etosha National Park, Namibia.
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Tony Heald / Lion (Panthera leo) male, walking amongst Devil's-thorn yellow flowers (Tribulus terrestris) Etosha National Park, Namibia.
Dubbeltjie flowers (Tribulus terrestris) with Lion (Panthera leo). Kgalagadi Transfrontier Park, South Africa, February.
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Richard Du Toit / Dubbeltjie flowers (Tribulus terrestris) with Lion (Panthera leo). Kgalagadi Transfrontier Park, South Africa, February.
White stork (Ciconia ciconia)  in grassland, Mapungubwe National Park, South Africa.
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Neil Aldridge / White stork (Ciconia ciconia) in grassland, Mapungubwe National Park, South Africa.
Ground squirrel (Xerus inauris) feeding in Devil's thorn flowers (Tribulus terrestris) Kgalagadi Transfrontier Park, Northern Cape, South Africa. Non-ex.
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Ann & Steve Toon / Ground squirrel (Xerus inauris) feeding in Devil's thorn flowers (Tribulus terrestris) Kgalagadi Transfrontier Park, Northern Cape, South Africa. Non-ex.
Ground squirrel (Xerus inauris) eating devil's thorn flowers (Tribulus zeyheri), Kgalagadi Transfrontier Park, Northern Cape, South Africa, February
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Ann & Steve Toon / Ground squirrel (Xerus inauris) eating devil's thorn flowers (Tribulus zeyheri), Kgalagadi Transfrontier Park, Northern Cape, South Africa, February
Lion (Panthera leo) in Dubbeltjie flower (Tribulus terrestris) Northern Cape Province, South Africa, February.
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Richard Du Toit / Lion (Panthera leo) in Dubbeltjie flower (Tribulus terrestris) Northern Cape Province, South Africa, February.
Yellow Vine (Tribulus terrestris) seed covered in sharp spines that enable dispersal, magnification 23x
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Albert Lleal / Yellow Vine (Tribulus terrestris) seed covered in sharp spines that enable dispersal, magnification 23x
Red Hartebeest (Alcelaphus caama) feeding on grasses near Yellow Vine (Tribulus terrestris) flowers, Kgalagadi Transfrontier Park, Botswana
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Richard Du Toit / Red Hartebeest (Alcelaphus caama) feeding on grasses near Yellow Vine (Tribulus terrestris) flowers, Kgalagadi Transfrontier Park, Botswana
Ostrich (Struthio camelus) male in Yellow Vine (Tribulus terrestris) flowers, Kgalagadi Transfrontier Park, Botswana
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Richard Du Toit / Ostrich (Struthio camelus) male in Yellow Vine (Tribulus terrestris) flowers, Kgalagadi Transfrontier Park, Botswana
RF- Zebra (Equus quagga) foal in profile standing in Devil's Thorn flowers (Tribulus terrestris). Etosha National Park, Namibia, January. (This image may be licensed either as rights managed or royalt...
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Tony Heald / RF- Zebra (Equus quagga) foal in profile standing in Devil's Thorn flowers (Tribulus terrestris). Etosha National Park, Namibia, January. (This image may be licensed either as rights managed or royalt...
Black-backed Jackal (Canis mesomelas) stalking through Devil's Thorn flowers (Tribulus terrestris). Etosha National Park, Namibia, January.
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Tony Heald / Black-backed Jackal (Canis mesomelas) stalking through Devil's Thorn flowers (Tribulus terrestris). Etosha National Park, Namibia, January.
White Rhinoceros (Ceratotherium simum) walking through Devil's Thorn flowers (Tribulus terrestris). Etosha National Park, Namibia, January.
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Sharon Heald / White Rhinoceros (Ceratotherium simum) walking through Devil's Thorn flowers (Tribulus terrestris). Etosha National Park, Namibia, January.
Crowned Plover (Vanellus coronatus) stood among Devil's Thorn flowers (Tribulus terrestris). Etosha National Park, Namibia, January.
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Tony Heald / Crowned Plover (Vanellus coronatus) stood among Devil's Thorn flowers (Tribulus terrestris). Etosha National Park, Namibia, January.
Greater kudu (Tragelaphus strepsiceros) male walking through Devil's Thorn flowers (Tribulus terrestris). Etosha National Park, Namibia, January.
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Sharon Heald / Greater kudu (Tragelaphus strepsiceros) male walking through Devil's Thorn flowers (Tribulus terrestris). Etosha National Park, Namibia, January.
Black-faced Impala (Aepyceros melamis petersi) male jumping ("stotting" or "pronking") to demonstrate fitness to potential mates, competitors and predators. Devil's Thorn fl (Tribulus terrestris) in f...
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Tony Heald / Black-faced Impala (Aepyceros melamis petersi) male jumping ("stotting" or "pronking") to demonstrate fitness to potential mates, competitors and predators. Devil's Thorn fl (Tribulus terrestris) in f...
Pin-Tailed Whydah (Vidua macroura) male in breeding plumage. Devil's Thorn flowers (Tribulus terrestris) cover the ground. Etosha National Park, Namibia, January.
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Tony Heald / Pin-Tailed Whydah (Vidua macroura) male in breeding plumage. Devil's Thorn flowers (Tribulus terrestris) cover the ground. Etosha National Park, Namibia, January.
Caltrop (Tribulus omanense) bush in flower, Liwa, Abu Dhabi, United Arab Emirates.
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Hanne & Jens Eriksen / Caltrop (Tribulus omanense) bush in flower, Liwa, Abu Dhabi, United Arab Emirates.
Caltrop (Tribulus omanense). Liwa, Abu Dhabi, United Arab Emirates.
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Hanne & Jens Eriksen / Caltrop (Tribulus omanense). Liwa, Abu Dhabi, United Arab Emirates.
Devil thorn flowers (Tribulus terrestris) after the rains, Kgalagadi Transfrontier Park, Kalahari desert, South Africa
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Jouan & Rius / Devil thorn flowers (Tribulus terrestris) after the rains, Kgalagadi Transfrontier Park, Kalahari desert, South Africa
Carpet of Devil thorn flowers (Tribulus terrestris) and other flowers during the rainy season, Namaqualand, South Africa
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Jouan & Rius / Carpet of Devil thorn flowers (Tribulus terrestris) and other flowers during the rainy season, Namaqualand, South Africa
Carpet of Devil thorn flowers (Tribulus terrestris) and other flowers during the rainy season, Namaqualand, South Africa
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Jouan & Rius / Carpet of Devil thorn flowers (Tribulus terrestris) and other flowers during the rainy season, Namaqualand, South Africa
Spitzkoppe, Namibia, with Devil's thorn {Tribulus sp} flowering in the foreground in the wet season
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Martin Gabriel / Spitzkoppe, Namibia, with Devil's thorn {Tribulus sp} flowering in the foreground in the wet season
Yellow Vine (Tribulus terrestris), Kgalagadi Transfrontier Park, South Africa
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Richard Du Toit / Yellow Vine (Tribulus terrestris), Kgalagadi Transfrontier Park, South Africa
Devil's claw (Harpagophytum procumbens) and Devil's thorn (Tribulus zeyheri), prominent creepers of the Kalahari Desert that appear after the rains, Kgalagadi Transfrontier Park, South Africa
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Philippe Clement / Devil's claw (Harpagophytum procumbens) and Devil's thorn (Tribulus zeyheri), prominent creepers of the Kalahari Desert that appear after the rains, Kgalagadi Transfrontier Park, South Africa
Devil's claw (Harpagophytum procumbens) and Devil's thorn (Tribulus zeyheri), prominent creepers of the Kalahari Desert that appear after the rains, Kgalagadi Transfrontier Park, South Africa
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Philippe Clement / Devil's claw (Harpagophytum procumbens) and Devil's thorn (Tribulus zeyheri), prominent creepers of the Kalahari Desert that appear after the rains, Kgalagadi Transfrontier Park, South Africa
Devil's thorn in flower {Tribulus terrestris} Sossusvlei, Namib desert, Namibia
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Pete Oxford / Devil's thorn in flower {Tribulus terrestris} Sossusvlei, Namib desert, Namibia
Kori bustard {Choriotis kori} amongst Little devil flowers {Tribulus terrestris} Kalahari Gemsbok NP, South Africa
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Richard Du Toit / Kori bustard {Choriotis kori} amongst Little devil flowers {Tribulus terrestris} Kalahari Gemsbok NP, South Africa
Waved Albatross (Phoebastria irrorata) pair bonding in flowering tribulus field during rains, Punta Cevallos, Espanola Island, Galapagos Islands, Ecuador
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Tui De Roy / Waved Albatross (Phoebastria irrorata) pair bonding in flowering tribulus field during rains, Punta Cevallos, Espanola Island, Galapagos Islands, Ecuador
Suricate (meerkat) {Suricata suricatta} on guard in flowering 'little devils' {Tribulus terrestris}. S Africa Kalahari >>> Gemsbok NP
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Richard Du Toit / Suricate (meerkat) {Suricata suricatta} on guard in flowering 'little devils' {Tribulus terrestris}. S Africa Kalahari >>> Gemsbok NP
Devil's thorn in flower {Tribulus terrestris} Clay castles, Skeleton coast, Namibia
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Pete Oxford / Devil's thorn in flower {Tribulus terrestris} Clay castles, Skeleton coast, Namibia
Desert in bloom. Devil thorn flowers {Tribulus terrestris} Kgalagadi TP, (formerely Kalahari Gemsbok) South Africa
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Pete Oxford / Desert in bloom. Devil thorn flowers {Tribulus terrestris} Kgalagadi TP, (formerely Kalahari Gemsbok) South Africa
Suricate (meerkat) {Suricata suricatta} on guard in flowering 'little devils' {Tribulus terrestris}. S Africa Kalahari >>> Gemsbok NP
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Richard Du Toit / Suricate (meerkat) {Suricata suricatta} on guard in flowering 'little devils' {Tribulus terrestris}. S Africa Kalahari >>> Gemsbok NP
Black backed jackal, with Flowering devils flowers {Tribulus terrestris} Kruger National Park, S. Africa
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Richard Du Toit / Black backed jackal, with Flowering devils flowers {Tribulus terrestris} Kruger National Park, S. Africa
Little devil flowers (Tribulus terrestris) and acacia Grey camelthorn trees, S Africa Kalahari Gemsbok NP in summer.
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Richard Du Toit / Little devil flowers (Tribulus terrestris) and acacia Grey camelthorn trees, S Africa Kalahari Gemsbok NP in summer.
Striped Ground Squirrel (Xerus erythropus) feeding. South Africa. Kalahari Gemsbok NP summer. Little Devil Flowers (Tribulus terrestris)
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Richard Du Toit / Striped Ground Squirrel (Xerus erythropus) feeding. South Africa. Kalahari Gemsbok NP summer. Little Devil Flowers (Tribulus terrestris)
Devil thorn bush {Tribulus terrestris} flowering in sand Skeleton coast, Namibia
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Pete Oxford / Devil thorn bush {Tribulus terrestris} flowering in sand Skeleton coast, Namibia
Devil thorn bush {Tribulus terrestris} in flower with seal bones in sand nearby, Skeleton coast, Namibia
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Pete Oxford / Devil thorn bush {Tribulus terrestris} in flower with seal bones in sand nearby, Skeleton coast, Namibia
African Lion (Panthera leo) female lying in Yellow Vine (Tribulus terrestris) flowers, Kgalagadi Transfrontier Park, South Africa
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Richard Du Toit / African Lion (Panthera leo) female lying in Yellow Vine (Tribulus terrestris) flowers, Kgalagadi Transfrontier Park, South Africa
Cape Ground Squirrel (Xerus inauris) feeding on Yellow Vine (Tribulus terrestris) flowers, Kgalagadi Transfrontier Park, Botswana
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Richard Du Toit / Cape Ground Squirrel (Xerus inauris) feeding on Yellow Vine (Tribulus terrestris) flowers, Kgalagadi Transfrontier Park, Botswana
Gemsbok (Oryx gazella) and Yellow Vine (Tribulus terrestris) flowers, Kgalagadi Transfrontier Park, Botswana
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Richard Du Toit / Gemsbok (Oryx gazella) and Yellow Vine (Tribulus terrestris) flowers, Kgalagadi Transfrontier Park, Botswana
African Lion (Panthera leo) male standing in Yellow Vine (Tribulus terrestris) flowers, Kgalagadi Transfrontier Park, South Africa
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Richard Du Toit / African Lion (Panthera leo) male standing in Yellow Vine (Tribulus terrestris) flowers, Kgalagadi Transfrontier Park, South Africa
Devil thorn bush {Tribulus terrestris} flowering in sand Skeleton coast, Namibia
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Pete Oxford / Devil thorn bush {Tribulus terrestris} flowering in sand Skeleton coast, Namibia
Female Cheetah (Acinonyx jubatus) with two juveniles, sitting among Devil's thorn (Tribulus terrestris) wildlfowers, Kgalagadi Transfrontier Park, Northern Cape, South Africa.
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Ann & Steve Toon / Female Cheetah (Acinonyx jubatus) with two juveniles, sitting among Devil's thorn (Tribulus terrestris) wildlfowers, Kgalagadi Transfrontier Park, Northern Cape, South Africa.
Female Cheetah (Acinonyx jubatus) lying down among Devil's thorn (Tribulus terrestris) flowers, Kgalagadi Transfrontier Park, Northern Cape, South Africa.
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Ann & Steve Toon / Female Cheetah (Acinonyx jubatus) lying down among Devil's thorn (Tribulus terrestris) flowers, Kgalagadi Transfrontier Park, Northern Cape, South Africa.