Bence Mate / Grey heron (Ardea cinerea) and Great egret (Ardea alba) fighting over fish, Lake Csaj, Pusztaszer, Hungary, December. Winner of the Portfolio category of the Terre Sauvage Nature Images Awards competi...
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Krijn Trimbos / Great crested grebe (Podiceps cristatus) two in territorial fight, The Netherlands, March.
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Espen Bergersen / Kittiwakes (Rissa tridactyla) squabbling on ice. Svalbard, Norway. July
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Ole Jorgen Liodden / Waved albatross (Phoebastria irrorata) pair in courtship display, Espanola, Galapagos Islands. Critically endangered species.
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Ole Jorgen Liodden / Meadow pipit (Anthus pratensis) feeding young Eurasian cuckoo (Cuculus canorus), Valdres, Norway, July.
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Roy Mangersnes / Polar bears (Ursus maritimus) courting on ocean ice north of Spitsbergen, Svalbard, Norway, July.
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Sam Hobson / Peregrine falcons (Falco peregrinus), juvenile female and male in flight, fighting over prey. Bristol, UK. June.
Rolf Nussbaumer / RF- Vermilion Flycatcher (Pyrocephalus rubinus) male giving insect to female for young in nest. Lake Corpus Christi, Texas, USA. May. (This image may be licensed either as rights managed or royalty fr...
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Konrad Wothe / Whooper Swan (Cygnus cygnus) pair arguing at wintering site, Kussharo-ko, Hokkaido, Japan
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Thomas Mangelsen / Bald Eagle (Haliaeetus leucocephalus) adults on snags, North America
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Michael Quinton / Trumpeter Swan (Cygnus buccinator) parent and squabbling day-old cygnets, North America
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Mitsuaki Iwago / Australian Sea Lion (Neophoca cinerea) male approaching a female to mate, Australia
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Andy Rouse / Adelie Penguin (Pygoscelis adeliae) squabbling on ice Antarctic Sound , Antarctica
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Andy Rouse / Adelie Penguin (Pygoscelis adeliae) squabbling on ice Antarctic Sound , Antarctica
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Roger Powell / Two Whiskered terns (Chlidonias hybridus) fighting over perch, Le Cherine Nature Reserve, La Brenne, France. April.
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Roger Powell / Two Whiskered terns (Chlidonias hybridus) fighting over perch, Le Cherine Nature Reserve, La Brenne, France. April.
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Roger Powell / Two Whiskered terns (Chlidonias hybridus) fighting over perch, Le Cherine Nature Reserve, La Brenne, France. April.
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Roger Powell / Two Whiskered terns (Chlidonias hybridus) disputing a perch on top of a post in a lake. Le Cherine Nature Reserve, La Brenne, France.
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David Kjaer / Griffon vulture (Gyps fulvus) flock on hillside with one interacting with Raven (Corvus corax) Pyrenees, Spain, April.
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Phil Savoie / Tree swallows (Tachycineta bicolor), perched on wire, calling aggressively to each other, Madison River, Montana, USA, June.
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Danny Green / RF - Polar Bear (Ursus maritimus) males fighting, Churchill, Canada, November (This image may be licensed either as rights managed or royalty free.)
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Richard Robinson / Snares crested penguin (Eudyptes robustus) pair arguing, Snares (Tine Heke) Sub Antarctic Island Group, Southern Ocean, New Zealand. Editorial use only.
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Ann & Steve Toon / Lappetfaced vulture (Torgos tracheliotos) harrassing whitebacked vulture (Gyps africanus) at carcass, Zimanga Private Game Reserve, South Africa, January.
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LYNN M. STONE / RF - Royal tern (Thalasseus maxima) offering fish to female as part of courtship. Mullet Key, St Petersburg. Florida, USA. (This image may be licensed either as rights managed or royalty free.)
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Danny Green / RF- Two Gannets (Morus) with nesting material. Great Saltee, County Wexford, Republic of Ireland. April . (This image may be licensed either as rights managed or royalty free.)
Roger Powell / Two rainbow lorikeets (Trichoglossus moluccanus) arguing on a branch. Werribee, Victoria, Australia.
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Carol Walker / Two wild Mustang stallions rearing in McCullough Peaks, Wyoming, USA. June.
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David Tipling / RF - King penguin (Aptenodytes patagonicus) and aggressive Antarctic fur seal pup (Arctocephalus gazella). St Andrews Bay, South Georgia. January. (This image may be licensed either as rights managed...
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David Tipling / RF - King penguins (Aptenodytes patagonicus) and aggressive Antarctic fur seal pup (Arctocephalus gazella). St Andrews Bay, South Georgia. January. (This image may be licensed either as rights managed...
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David Tipling / RF - Antarctic fur seal (Arctocephalus gazella) females fighting. Albatross Island, South Georgia. January. (This image may be licensed either as rights managed or royalty free.)
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David Tipling / RF - Wandering albatross (Diomedea exulans) pair displaying at dusk. Albatross Island, South Georgia. January. (This image may be licensed either as rights managed or royalty free.)
Edwin Giesbers / Red foxes (Vulpes vulpes) fighting.
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Danny Green / Common cranes (Grus grus) fighting, Hula Valley, Northern Israel, January
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Gabriel Rojo / Bay-winged cowbird (Agelaioides badius) two perched on branch in rain, one calling and another feeding, Calden forest, La Pampa, Argentina
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Espen Bergersen / Polar bear (Ursus maritimus) aggression, two roaring at each other measuring strength, Spitsbergen, Svalbard, Norway. July. GPS collar digitally removed.
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Andy Rouse / RF- Mountain hares (Lepus timidus) boxing in snow, Scotland, UK, December. (This image may be licensed either as rights managed or royalty free.)
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Frederique Olivier / Adelie penguin (Pygoscelis adeliae) fighting with neighbour with chick between, Antarctica.
XI ZHINONG / Gentoo penguins (Pygoscelis papua) 'arguing' Antarctica, November.
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Andrew Parkinson / Gannet (Morus bassanu) two breeding pairs 'arguing' on a coastal nest site. Shetland Islands, Scotland, UK, July.
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Andy Trowbridge / Canada geese (Branta canadensis) on frozen lake 'arguing', Southern Norway, March.
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Denis-Huot / RF- African Elephant (Loxodonta africana) males fighting. Masai-Mara Game Reserve, Kenya. (This image may be licensed either as rights managed or royalty free.)
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Jabruson / African Forest elephant (Loxodonta africana cyclotis) two bulls sparring, competition for access to valuable minerals located at certain points within the clearing's water points, Dzanga-Ndoki Nationa...
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Tui De Roy / Waved Albatross (Phoebastria irrorata) and Blue-footed Booby (Sula nebouxii) squabbling over nesting sites. Galapagos Islands, Ecuador, June.
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Edwin Giesbers / RF- Alpine marmots (Marmota marmota) fighting, Hohe Tauern National Park, Austria. July.
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Andy Rouse / RF- Grizzly Bear (Ursus arctos horribilis) male (right) and female fighting in water over salmon. Katmai, Alaska, USA, August. (This image may be licensed either as rights managed or royalty free.)
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Edwin Giesbers / RF- Two Kodiak / Alaskan brown bears (Ursus arctos middendorffi) fighting, captive. (This image may be licensed either as rights managed or royalty free.)
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Andy Rouse / RF- Asian Elephant (Elaphas maximus) with Black winged stilt (Himantopus himantopus) showing aggression, Minneriya National Park, Sri Lanka. Endangered species. (This image may be licensed either as r...
Duncan Usher / Hawfinch (Coccothraustes coccothraustes) and Eurasian Tree Sparrow (Passer montanus) quarreling, Lower Saxony, Germany
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Konrad Wothe / Steller's Sea Lion (Eumetopias jubatus) group hauled out on rocks, Inside Passage, Alaska
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Jan Vermeer / Gentoo Penguin (Pygoscelis papua) pair arguing in the snow, Antarctica
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Konrad Wothe / Black-tailed Prairie Dog (Cynomys ludovicianus) young tugging at either end of a blade of grass, Arizona
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Konrad Wothe / Northern Elephant Seal (Mirounga angustirostris) juvenile males play fighting, North America
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Richard Du Toit / Pied Kingfisher (Ceryle rudis) pair perching, Chobe River, Caprivi Strip, Namibia
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Jose B. Ruiz / Common crane {Grus grus} arguing, Laguna de Gallocanta, Teruel, Aragón, Spain
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Pete Oxford / Hood Mockingbird (Nesomimus macdonaldi) pair arguing, vulnerable, Espanola Island, Galapagos Islands, Ecuador
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Pete Oxford / Royal Penguins {Eudyptes schlegeli} arguing, Sandy Bay, Macquarie Island, Tasmania, Australia
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Staffan Widstrand / RF- Polar bears play fighting (Ursus maritimus). Churchill, Manitoba, Canada. (This image may be licensed either as rights managed or royalty free.)
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Matthias Breiter / Grizzly Bear (Ursus arctos horribilis) mother defending her offspring from another bear, Katmai National Park, Alaska
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Duncan Usher / Marsh Tit (Parus palustris) and Great Tit (Parus major) fighting over Sunflower Seeds, Europe
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Flip de Nooyer / Southern Elephant Seal (Mirounga leonina) pair of sparring pups in shallow water, South Georgia Island
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Duncan Usher / European Greenfinch (Chloris chloris) and Great Tit (Parus major) arguing over sunflower seeds, Europe
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Tim Fitzharris / African Fish Eagle (Haliaeetus vocifer), quarreling with Marabou Stork (Leptoptilos crumeniferus), Kenya
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Konrad Wothe / Whooper Swan (Cygnus cygnus) pair wintering, Hokkaido, Japan
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Thomas Mangelsen / Polar Bear (Ursus maritimus) pair of males fighting, Churchill, Manitoba, Canada
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Tim Fitzharris / Steller's Sea Lion (Eumetopias jubatus) pair, Brothers Island, Alaska
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Konrad Wothe / Hyacinth Macaw (Anodorhynchus hyacinthinus) pair perching, Pantanal, Brazil
David Tipling / Two Blue tits (Cyanistes caeruleus) fighting in garden, Norfolk, UK. February.
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Tui De Roy / Two Floreana giant tortoises hybrid descendant (Chelonoidis elephantopus) in confrontation, National Park Breeding Center, Santa Cruz Island, Galapagos Islands. Captive.
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Ann & Steve Toon / Two Lions (Panthera leo) male, arguing over young Wildebeest prey (Connochaetes taurinus), Kgalagadi Transfrontier Park, Northern Cape, South Africa.
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TOM MANGELSEN / Antarctic fur seal (Arctocephalus gazella) and King penguin (Aptenodytes patagonicus) squabbling, face to face, on beach, Salisbury Plain, South Georgia, South Atlantic Ocean.
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Robin Chittenden / Black-headed gull (Chroicocephalus ridibundus) in confrontation with Coot (Fulica atrica) whilst swimming, Whitlingham Country Park, Norfolk, UK. December.
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Staffan Widstrand / Four Black vultures (Coragyps atratus) fighting over a Caiman (Caiman yacare) carcass in river, Cuiaba River, Pantanal wetlands, Mato Grosso, Brazil.
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Enrique Lopez-Tapia / Buddhist monks in Dialectics Courtyard, where monks debate and discuss religious topics, in Sera Monastery, founded in 1419. Tibet, China.
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Gabriel Rojo / Two Giant petrels (Macronectes giganteus) bickering on beach, Peninsula Valdes, Patagonia, Argentina.
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Jussi Murtosaari / Two Eurasian coot (Fulica atra) fighting over territory quarrel, Finland. April.
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Bernard Castelein / Two male Village weaver (Ploceus cucullatus) perched in tree, squabbling, Allahein river, The Gambia.
