Marion Vollborn/ BIA / Whooper Swan (Cygnus cygnus) group taking flight on snow, Hokkaido, Japan

Tim Fitzharris / Trumpeter Swan (Cygnus buccinator) flock flying, Arkansas

Donald M. Jones / Trumpeter Swan (Cygnus buccinator) trio flying, southern Alaska

Michel Roggo / Chub fish (Leuciscus cephalus) and Mute swan (Cygnus color) swimming over, Rhine river, Schaffhausen, Switzerland. RUNNER-UP - Wildlife Photographer of the Year 2010.

Michael Quinton / Trumpeter Swan (Cygnus buccinator) pair on lake at sunset, North America

Flip de Nooyer / Mute Swan (Cygnus olor) three cygnets on parent's back, Europe

Flip de Nooyer / Mute Swan (Cygnus olor) pair courting, Europe

Konrad Wothe / Whooper Swan (Cygnus cygnus) pair arguing at wintering site, Kussharo-ko, Hokkaido, Japan

Konrad Wothe / Mute Swan (Cygnus olor) with three chicks riding on her back, Germany

Konrad Wothe / Mute Swan (Cygnus olor) courting pair, Germany

Jim Brandenburg / Tundra Swan (Cygnus columbianus) trio flying against cloudy sky, Minnesota

Michael Quinton / Trumpeter Swan (Cygnus buccinator) pair on lake, Copper River Delta, Alaska

Jim Brandenburg / Tundra Swan (Cygnus columbianus) flock taking off from lake, Mattamuskeet National Wildlife Reserve, North Carolina

Tui De Roy / Black-necked Swan (Cygnus melancoryphus) group swimming, Puerto Natales, Chilean Fjords, Chile

Michael Quinton / Trumpeter Swan (Cygnus buccinator) pair on lake, North America

Michael Quinton / Trumpeter Swan (Cygnus buccinator) parent and squabbling day-old cygnets, North America

Martin Grimm/ BIA / Mute Swan (Cygnus olor), Baden-Wurttemberg, Germany

Jeff Foott / Trumpeter Swan (Cygnus buccinator) parents and cygnets, National Elk Refuge, Wyoming

Donald M. Jones / Trumpeter Swan (Cygnus buccinator) parents and juveniles preening, Montana

Chase Dekker / Trumpeter Swan (Cygnus buccinator) in misty river, Grand Teton National Park, Wyoming

Marion Vollborn/ BIA / Whooper Swan (Cygnus cygnus) taking flight on snow, Hokkaido, Japan

Marion Vollborn/ BIA / Whooper Swan (Cygnus cygnus) flying, Hokkaido, Japan

Sean Crane / Trumpeter Swan (Cygnus buccinator) pair on lake in autumn, Tern Lake, Alaska

Holger Duty/ BIA / Mute Swan (Cygnus olor) on nest, Mecklenburg-Vorpommern, Germany

Folkert Christoffers/ BIA / Mute Swan (Cygnus olor) on nest, Lower Saxony, Germany

Holger Doernhoff/ BIA / Black Swan (Cygnus atratus), North Rhine-Westphalia, Germany

Agustin Esmoris / Black-necked Swan (Cygnus melancoryphus), Argentina

Ralf Kistowski/ BIA / Black Swan (Cygnus atratus), North Rhine-Westphalia, Germany

Rosl Roessner/ BIA / Mute Swan (Cygnus olor), Bavaria, Germany

Mark Carwardine / Mute Swan (Cygnus olor), Slimbridge, Gloucestershire, England, United Kingdom

Mark Carwardine / Tundra Swan (Cygnus columbianus) calling, Slimbridge, Gloucestershire, England, United Kingdom

Jim Brandenburg / Trumpeter Swan (Cygnus buccinator) pair dabbling, Minnesota

Hans Glader/ BIA / Whooper Swan (Cygnus cygnus) group in lake in winter, Finland

Hans Glader/ BIA / Whooper Swan (Cygnus cygnus) group in lake in winter, Finland

Hans Glader/ BIA / Whooper Swan (Cygnus cygnus) group in lake in winter, Finland

Marion Vollborn/ BIA / Whooper Swan (Cygnus cygnus) group taking flight, Hokkaido, Japan

Jan Vermeer / Whooper Swan (Cygnus cygnus), Lake Kussharo, Akan National Park, Japan

Jan Vermeer / Whooper Swan (Cygnus cygnus) on ice, Lake Kussharo, Akan National Park, Japan

Jan Vermeer / Whooper Swan (Cygnus cygnus) group in snowstorm, Lake Kussharo, Akan National Park, Japan

Jan Vermeer / Whooper Swan (Cygnus cygnus) pair flying in snowstorm, Lake Kussharo, Akan National Park, Japan

Jan Vermeer / Whooper Swan (Cygnus cygnus) trio landing, Lake Kussharo, Akan National Park, Japan

Jan Vermeer / Whooper Swan (Cygnus cygnus) group flying in snowstorm, Lake Kussharo, Akan National Park, Japan

Tim Fitzharris / Trumpeter Swan (Cygnus buccinator) pair, Magness Lake, Arkansas

Tim Fitzharris / Trumpeter Swan (Cygnus buccinator) group flying, Magness Lake, Arkansas

Konrad Wothe / Mute Swan (Cygnus olor) parent with cygnets, Bavaria, Germany

Donald M. Jones / Tundra Swan (Cygnus columbianus) flock flying, central Montana

Donald M. Jones / Tundra Swan (Cygnus columbianus) flock flying near windmill, central Montana

Donald M. Jones / Tundra Swan (Cygnus columbianus) flock flying, central Montana

Donald M. Jones / Tundra Swan (Cygnus columbianus) pair flying, central Montana

Donald M. Jones / Tundra Swan (Cygnus columbianus) flying, central Montana

Donald M. Jones / Tundra Swan (Cygnus columbianus) flying, central Montana

Donald M. Jones / Trumpeter Swan (Cygnus buccinator) parent with cygnet, western Montana

Donald M. Jones / Trumpeter Swan (Cygnus buccinator) parent with cygnet, western Montana

Donald M. Jones / Trumpeter Swan (Cygnus buccinator) parent with cygnets, western Montana

Donald M. Jones / Trumpeter Swan (Cygnus buccinator) calling, southern Alaska

Donald M. Jones / Trumpeter Swan (Cygnus buccinator) trio flying, southern Alaska

Donald M. Jones / Trumpeter Swan (Cygnus buccinator) flying, southern Alaska

Donald M. Jones / Trumpeter Swan (Cygnus buccinator) trio swimming, southern Alaska

Donald M. Jones / Trumpeter Swan (Cygnus buccinator) pair fighting, southern Alaska

Donald M. Jones / Trumpeter Swan (Cygnus buccinator) taking flight, southern Alaska

Donald M. Jones / Trumpeter Swan (Cygnus buccinator) on nest, southern Alaska

Tim Fitzharris / Trumpeter Swan (Cygnus buccinator) flapping, Magness Lake, Arkansas

Tim Fitzharris / Trumpeter Swan (Cygnus buccinator) trio flying, Magness Lake, Arkansas

Tim Fitzharris / Trumpeter Swan (Cygnus buccinator) trio landing, Magness Lake, Arkansas

Tim Fitzharris / Trumpeter Swan (Cygnus buccinator) flapping, Magness Lake, Arkansas

Tim Fitzharris / Trumpeter Swan (Cygnus buccinator) trio flying, Magness Lake, Arkansas

Tim Fitzharris / Trumpeter Swan (Cygnus buccinator) pair, Magness Lake, Arkansas

Tim Fitzharris / Trumpeter Swan (Cygnus buccinator) pair preening, Magness Lake, Arkansas

Donald M. Jones / Trumpeter Swan (Cygnus buccinator), western Montana

Tim Fitzharris / Trumpeter Swan (Cygnus buccinator) on lake, Boxley Valley, Buffalo National River, Arkansas

Peter Waechtershaeuser / Whooper Swan (Cygnus cygnus) flock in lake in winter, Hokkaido, Japan

Duncan Usher / Mute Swan (Cygnus olor) parents leading cygnets to lake through a park, Germany

Peter Waechtershaeuser / Whooper Swan (Cygnus cygnus) group swimming along shore, Hokkaido, Japan

Jan Wegener / Black Swan (Cygnus atratus), Victoria, Australia

Rob Drummond / Black Swan (Cygnus atratus), Victoria, Australia

Rob Drummond / Black Swan (Cygnus atratus) flying, Victoria, Australia

Matthew Studebaker / Trumpeter Swan (Cygnus buccinator) pair, Michigan

Alan Murphy / Trumpeter Swan (Cygnus buccinator), British Columbia, Canada

Rob Drummond / Black Swan (Cygnus atratus), Victoria, Australia

Kevin Schafer / Black-necked Swan (Cygnus melancoryphus) pair, Patagonia, Chile

Thomas Marent / Mute Swan (Cygnus olor), Switzerland

Michael Quinton / Trumpeter Swan (Cygnus buccinator), Alaska

Michael Quinton / Trumpeter Swan (Cygnus buccinator) pair, Alaska

Cyril Ruoso / Mute Swan (Cygnus olor) male on Yonne, France

Cyril Ruoso / Mute Swan (Cygnus olor) female on Yonne, France

Cyril Ruoso / Whooper Swan (Cygnus cygnus) fighting, east Iceland

Cyril Ruoso / Whooper Swan (Cygnus cygnus) flying low over the water, east Iceland

Cyril Ruoso / Whooper Swan (Cygnus cygnus) male chasing intruder from nesting area, east Iceland

Cyril Ruoso / Whooper Swan (Cygnus cygnus), Iceland

Cyril Ruoso / Whooper Swan (Cygnus cygnus) portrait, Iceland

Cyril Ruoso / Whooper Swan (Cygnus cygnus), Iceland

Cyril Ruoso / Whooper Swan (Cygnus cygnus), Iceland

Cyril Ruoso / Whooper Swan (Cygnus cygnus) drinking, Iceland

Konrad Wothe / Mute Swan (Cygnus olor) in defensive posture, Upper Bavaria, Germany

Hiroya Minakuchi / Whooper Swan (Cygnus cygnus) group flying over Northern Pintail (Anas acuta) flock, Lake Hyo, Japan

Hiroya Minakuchi / Whooper Swan (Cygnus cygnus) and Northern Pintail (Anas acuta) flock, Lake Hyo, Japan