Ingo Arndt / Grizzly bear (Ursus arctos horribilis) silhouette, Lake Clark National Park, Alaska

Will Watson / Atlantic salmon (Salmo salar) leaping a weir, Shrewsbury, River Severn, Shropshire, England, UK. November.

Wim van den Heever / Vervet Monkey (Chlorocebus pygerythrus) baby jumping between branches, photographed mid air. Okavango Delta Botswana.

Wim van den Heever / Black heron (Egretta ardesiaca) fishing and using wings to create an area of shade to attract fish, on the banks of the Chobe River, Botswana.

Markus Varesvuo / Two male Black grouse (Tetrao / Lyrurus tetrix) fighting, Kuusamo, Finland, April.

Brent Stephenson / Young Polar bear (Ursus maritimus) walking across melting sea ice, Scott Inlet, Baffin Island, Canadian Arctic, August. Vulnerable species.

Wim van den Heever / Leopard (Panthera pardus) hunting Springbok (Antidorcas marsupialis) Etosha, Namibia, Finalist in the Mammals Category of the Wildlife Photographer of the Year 2015.

Doug Perrine / Humpback whale (Megaptera novaeangliae) lone female playing, blowing bubble stream, Kona, Hawaii, USA.

Sven Zacek / Ural owl (Strix uralensis) chick trying to climb an aspen tree with its beak, talons and wings, Southern Estonia. June.

Claudio Contreras / Caribbean flamingo (Phoenicopterus ruber) preparing to sleep, Ria Lagartos Biosphere Reserve, Yucatan Peninsula Mexico, July. Bookplate.

Bence Mate / Little egret (Egretta garzetta) shaking water off wings at sunset, Lake Csaj, Pusztaszer, Hungary. May. Winner of the Portfolio category of the Terre Sauvage Nature Images Awards competition 2015.

Bence Mate / Grey heron (Ardea cinerea) and Great egret (Ardea alba) fighting over fish, Lake Csaj, Pusztaszer, Hungary, December. Winner of the Portfolio category of the Terre Sauvage Nature Images Awards competi...

Bence Mate / Two Great egrets (Ardea alba) standing opposite each other with Grey heron (Ardea cinerea) in between. Lake Csaj, Pusztaszer, Hungary, January. Winner of the Portfolio category of the Terre Sauvage Na...

Alex Mustard / Portrait of Bigfin squid (Sepioteuthis lessoniana) hovering in open water above a coral reef at night. Dampier Strait, Raja Ampat, West Papua, Indonesia. Tropical West Pacific Ocean

Alex Mustard / Galapagos sea lions (Zalophus wollebaeki) young playing in shallow water. Floreana Island, Galapagos Islands, Ecuador. East Pacific Ocean

Charlie Summers / Rocky mountain bighorn (Ovis canadensis) ram jumping a barbed wire fence, Montana, USA,

Dave Watts / Northern bald ibis (Geronticus eremita) feeding, Morocco, Critically endangered species.

Charlie Summers / Adelie Penguin (Pygoscelis adeliae) carrying stolen rock from another nest in rookery, Peterman Island, Antarctica, November.

Tony Heald / Junglefowl (Gallus gallus) male, Kaziranga National Park, Assam, India, March.

Sharon Heald / Hippopotamus (Hippopotamus amphibius) charging, Chobe National Park, Botswana, July.

Tony Heald / Hanuman langur (Semnopithecus entellus) eating coconut husk, Bandhavgarh National Park, India. March.

Juan Carlos Munoz / Northern tamandua (Tamandua mexicana) climbing tree, Corcovado National Park, Osa Peninsula, Puntarenas Province, Costa Rica.

Hermann Brehm / Reindeer (Rangifer tarandus) in woodland, Sweden, September.

Arne Hodalic / Nenet herdsman with Reindeer (Rangifer tarandus) during summer migration, Yamal Peninsula, Russia. May.

Pete Oxford / Gayal / Mithun (Bos frontalis) on river bank. Arunachal Pradesh, North East India. November 2014.

Uri Golman / Greenland reindeer (Rangifer tarandus groenlandicus) in velvet, Kangerlussuaq, Greenland, June.

Uri Golman / Forest reindeer, (Rangifer tarandus fennicus) in velvet, Viiksimo, Kuhmo region. Finland, July. Rare subspecies which were nearly extinct in the 19th Century.

Richard Kirby / Svalbard reindeer (Rangifer tarandus platyrhynchus) mother and calves, Svalbard, Norway, February.

Cyril Ruoso / Sumatran orangutan (Pongo abelii) female holding hands with baby whilst hanging from liana. Gunung Leuser National Park, Sumatra, Indonesia.

Pal Hermansen / European moose (Alces alces) bulls near houses, Nordland, Norway. July.

Will Burrard-Lucas / Ethiopian wolf (Canis simensis) stretching as it wakes up, Web Valley, Ethiopia.

Sven Zacek / American mink (Mustela vison) on ice with Common frog (Rana temporaria) prey, Tartumaa, Estonia, January. Introduced species.

Sven Zacek / Cross-country skiier on a moonlit night admiring the Northern Lights (Aurora borealis), Norway, March 2013.

Sven Zacek / Roe deer (Capreolus capreolus) buck with velvet antlers in snowy field at sunset, Tartumaa, Estonia, March.

Sven Zacek / White-tailed sea eagle (Haliaeetus albicilla) flying, and hunting over sea, Norway, July.

Yukihiro Fukuda / Japanese macaque (Macaca fuscata fuscata) rare white furred baby jumping, Jigokudani Valley, Nagano Prefecture, Japan. June.

Solvin Zankl / Deepsea Isopod (Glyptonotus antarcticus) from Antarctic Ocean at a depth of 231m.

Bruno D'Amicis / Fennec fox (Vulpes zerda) adult at night digging for prey among the roots of Retam broom shrub. Grand Erg Oriental, Kebili Governorate, Tunisia. Taken with remote camera trap.

George Sanker / Snowy egret (Egretta thula) with plumes erect, St. Augustine, Florida, USA, April.

George Sanker / Great egret (Ardea alba) taking off with fish prey in mouth, Myakka River State Park, Florida, USA, March.

George Sanker / Barred owl (Strix varia) stretching wings, Corkscrew Swamp Audubon Sanctuary, Florida, USA, March.

Richard Du Toit / Impala (Aepyceros melampus) leaping, silhouetted at dawn, Itala Game Reserve, KwaZulu-Natal, South Africa.

Richard Du Toit / Greater kudu (Tregalaphus strepsiceros) covered in Yellowbilled oxpeckers (Buphagus africanus) rear view, Kruger National Park, South Africa.

Dong Lei / Crocodile newt (Tylototriton verrucosus) Gaoligong Mountain National Nature Reserve, Tengchong county, Yunnan Province, China. May.

Andy Rouse / Great crested grebe (Podiceps cristatus cristatus) courtship dance at dawn, Cardiff, UK, March.

Anup Shah / African elephants (Loxodonta africana) wallowing at a waterhole, wide angle view. Maasai Mara National Reserve, Kenya.

Pete Oxford / Galapagos land iguana (Conolophus subcristatus) walking, South Plaza Island, Galapagos. Endemic.

Pete Oxford / Marine iguana (Amblyrhynchus cristatus) underwater. Fernandina Island. Galapagos, Endemic Species.

Staffan Widstrand / Arctic fox (Vulpes lagopus) in winter coat, sitting, Spitsbergen, Svalbard, Norway, September

Gabriel Rojo / Guanaco (Lama guanicoe) in front of mountainous landscape, Torres del Paine National Park, Chile

Espen Bergersen / Red fox (Vulpes vulpes) walking through snow, Lapland, Finland, March.

Danny Green / Coastal brown bear (Ursus arctos) standing on hind legs, Lake Clarke National Park, Alaska, September.

Danny Green / Whooper swans (Cygnus cygnus) two in flight, during snowfall, Lake Kussharo, Japan, February

Danny Green / Japanese cranes (Grus japonensis) displaying in snow, Hokkaido, Japan, February

Enrique Lopez-Tapia / Karo boys with decorative skin painting.Territory of the Karo tribe. Omo river. Ethiopia, November 2014

Franco Banfi / Pygmy blue whale (Balaenoptera musculus brevicauda) subspecies of blue whale, Mirissa, Sri Lanka, Indian Ocean. Endangered species.

Franco Banfi / Pygmy blue whale (Balaenoptera musculus brevicauda) subspecies of blue whale, diving down, Mirissa, Sri Lanka, Indian Ocean. Endangered species.

David Pattyn / Alpine ibex (Capra ibex) adult male in mountain landscape. Alps, Aosta Valley, Gran Paradiso National Park, Italy. September.

Ole Jorgen Liodden / Polar bear (Ursus maritimus) walking over blue ice, Nordaustlandet, Svalbard, Norway, July.

Ole Jorgen Liodden / Polar bear (Ursus maritimus) waving standing on hind legs with paws raised, Svalbard, Norway, September.

Ole Jorgen Liodden / Common sandpiper (Actitis hypoleucos) singing with tongue visible, Norway, June.

Doug Perrine / Mediterranean monk seal (Monachus monachus) adult male swimming, Deserta Grande, Ilhas Desertas, Parque Natural da Madeira, Portugal, Atlantic Ocean, June. Critically endangered species

Kristel Richard / Przewalski / Takhi Horse (Equus ferus przewalskii) breeding stallion stands alert, Hustai National Park, Tuv Province, Mongolia. Endangered species. September.

Andy Rouse / Bengal tiger (Panthera tigris tigris) 11 month cubs play fighting, Ranthambhore National Park, India.

Andy Rouse / Gentoo penguin (Pygoscelis papua) jumping onto beach, Carcass Island, Falkland Islands.

Steven Kazlowski / Polar bear (Ursus maritimus) juvenile crossing over newly forming pack ice, during autumn freeze up, Beaufort Sea, off Arctic coast, Alaska

Tui De Roy / Erect-crested penguin (Eudyptes sclateri) pair. Antipodes Island, New Zealand Sub-Antarctic Islands. Endemic to Antipodes and Bounty Islands. Endangered species.

Pete Oxford / White rhinoceros calf (Ceratotherium simum) Great Karoo. Private Reserve, South Africa. Endangered species

Dong Lei / Group of local children smiling, near Lantsang Mekong river, Kawakarpo Mountain, Meri Snow Mountain National Park, Yunnan Province, China.

Nick Garbutt / Male Bare-faced Curassow (Crax fasciolata) near Araras Lodge, Mato Grosso, Northern Pantanal, Brazil, South America.

Nick Garbutt / Zebu Cattle (Bos primigenius indicus), Los Llanos, Colombia, South America.

Pascal Kobeh / Marine iguana (Amblyrhynchus cristatus) swimming back to the surface, Galapagos. Pacific ocean.

Ben Cranke / White Woodpecker (Melanerpes candidus) perched on a tree branch, Piaui, Brazil. August.

Ben Cranke / Black-striped capuchin (Sapajus libidinosus) using rock as a tool to break open palm nut Parnaiba Headwaters National Park, Piaui, Brazil. July.

Ben Cranke / Black-striped capuchin (Sapajus libidinosus) using rock as a tool to break open palm nut, Parnaiba Headwaters National Park, Piaui, Brazil. July.

Mark Taylor / Playful tabby male kitten, age 10 weeks.

Mark Taylor / Bengal cat standing on a fallen tree.

Richard Robinson / William Trubridge diving - a world champion free diver who is campaigning to protect the Critically Endangered Maui's dolphin (Cephalorhynchus hectori maui) endemic to New Zealand. January 2008. Edi...

Neil Aldridge / Pair of Blesbok (Damaliscus pygargus) galloping across the open flood plains of the Bushman's River, Kariega Game Reserve, South Africa. Overall winner of the GDT Competition 2014.

Nick Garbutt / Adult male Diademed Sifaka (Propithecus diadema) 'skipping / dancing' across open ground between forest fragments, eastern Madagascar.

Olga Kamenskaya / Man looking into ice hole dug for ice diving, Lake Baikal, Siberia, Russia, March. Model released.

Andy Trowbridge / Female goshawk (Accipiter gentilis) hiding Wood pigeon (Columba palumbus) prey from passing crows. Southern Norway, February.

Andy Trowbridge / Arctic Fox (Alopex / Vulpes lagopus) at sunset, during moult from grey summer fur to winter white. Dovrefjell National Park, Norway, September.

David Pattyn / Hanuman langur / Northern Plains Grey Langur (Semnopithecus entellus) drinking from a water tap at Ranthambore fort. Ranthambore National Park, Rajasthan, India.

Jabruson / African rainforest clearance, men sawing hardwood tree trunks to make planks. South of Mbomo, Odzala-Kokoua National Park, Republic of Congo, May 2005.

Robin Chittenden / European robin (Erithacus rubecula) perched on branch, Norwich, Norfolk, UK, July.

Peter Cairns / Tourists watching American bison (Bison bison) Yellowstone National Park, Wyoming, USA, February 2013.

Inaki Relanzon / Zebu cattle (Bos indicus) market in Ambalavao, central Madagascar, November 2012.

Anup Shah / African elephants (Loxodonta africana) drinking at a waterhole. Masai Mara National Reserve, Kenya. Taken with remote wide angle camera.

Andy Rouse / Giant Panda (Ailuropoda melanoleuca) cub climbing tree. Chengdu, China. Captive.

Jabruson / 'Kith' / 'Kin' Sumatran orangutan (Pongo abelii) captive animal in walled enclosure, Perth Zoo, Western Australia, August 2001.

David Woodfall / Residents walking through February 2014 floods from River Thames, Wraysbury, Surrey, England, UK, 16th February 2014.

Jussi Murtosaari / European brown bear (Ursus arctos arctos) standing on one foot, northern Finland, May.

Eric Medard / Grey heron (Ardea cinerea) in small stream at night, taken with infra-red remote camera trap, France. October.

Eric Medard / European brown bear (Ursus arctos arctos) walking through forest at night, taken with infra red remote camera trap, Slovenia, October.

Eric Baccega / Male Ostrich (Struthio camelus) protecting chicks from the sun with its wings, Samburu National Reserve, Kenya, Africa.