Jose B. Ruiz / Sea daffodil and flying bee, coastal sand dunes, Spain

David Woodfall / Honey bee (Apis mellifera) on Sea holly (Eryngium) Wales, UK, July.

Andy Sands / Mining bee (Andrena pilipes) female carrying pollen returning to burrow on coastal cliff top, Cornwall, England, UK, September

Nick Upton / Moss carder bumblebee (Bombus muscorum) visiting Red apple Ice plant / Baby sunrose (Aptenia cordifolia) an invasive South African species flowering on a coastal headland, Vale de Figueira, Portugal,...

Nick Upton / Moss carder bumblebee (Bombus muscorum) visiting Red apple Ice plant / Baby sunrose (Aptenia cordifolia) an invasive South African species flowering on a coastal headland, Vale de Figueira, Portugal,...

MYN / Corne van der Linden / Solitary bee, (Hylaeus ruficeps kalamundae) female, Swan Coastal Plain, South Western Australia.

MYN / Corne van der Linden / Leafcutter bee (Megachile) female, Swan Coastal Plain, South Western Australia. December.

MYN / Corne van der Linden / Leafcutter bee (Megachile aurifrons) male, Swan Coastal Plain, South Western Australia. December.

MYN / Corne van der Linden / Leafcutter bee (Megachile aurifrons) female, Swan Coastal Plain, South Western Australia. December.

MYN / Corne van der Linden / Solitary bee (Lipotrichis sp) Swan Coastal Plain, South-Western Australia, November.

MYN / Gil Wizen / Cuckoo bee (Thyreus sp) a cleptoparasite of other species of bees, Central Coastal Plain, Israel, June. meetyourneighbours.net project

MYN / Gil Wizen / Cuckoo bee (Thyreus sp) a cleptoparasite of other species of bees, Central Coastal Plain, Israel, June. meetyourneighbours.net project

B&S Draker / Honey Bee (Apis mellifera) in flight on White Prickly Poppy (Argemone albiflora). Sinton, Corpus Christi, Coastal Bend, Texas, USA, June.

Mark Moffett / Orchid (Clusia lanceolata) a native to Brazil's Atlantic Coastal forest, the female Orchid Bee (Euglossa viridissima) shown here, gathers floral resins from the flower to construct her brood cells, At...

Premaphotos / Coast leaf-cutter bee (Megachile maritima) female feeding on Restharrow flower on coastal dunes, UK

Nick Upton / Long-horned bee (Eucera longicornis) male, nectaring from Greater bird's-foot trefoil (Lotus pedunculatus) flower on coastal grassland, The Lizard, Cornwall, England, UK. June.

Nick Upton / Long-horned bee (Eucera longicornis) male, flying towards Greater bird's-foot trefoil (Lotus pedunculatus) flowers on coastal grassland, The Lizard, Cornwall, England, UK. June.

Nick Upton / Long-horned bee (Eucera longicornis) female, emerging from its nest burrow in sandy coastal cliffs, The Lizard, Cornwall, England, UK. June.

Nick Upton / Long-horned bee (Eucera longicornis) female, emerging from its nest burrow in sandy coastal cliffs, The Lizard, Cornwall, England, UK. June.

Nick Upton / Honey bees (Apis mellifera) at entrance to a wild hive among tree roots in coastal woodland, South Cornwall, England, UK. June.

Nick Upton / Shrill carder bee (Bombus sylvarum) nectaring on Yellow rattle (Rhinanthus major) flowers on permanent coastal sand dune grassland, Kenfig National Nature Reserve, Glamorgan, Wales, UK. June.

Nick Upton / Red-tailed bumblebee (Bombus lapidarius) worker emerging from nest burrow in mature, stable sand dunes, Kenfig National Nature Reserve, Glamorgan, Wales, UK. June.

Nick Upton / Red-tailed bumblebee (Bombus lapidarius) worker taking off from entrance of nest burrow in mature, stable sand dunes, Kenfig National Nature Reserve, Glamorgan, Wales, UK. June.

Nick Upton / Red-tailed bumblebee (Bombus lapidarius) worker entering its nest burrow in mature, stable sand dunes, Kenfig National Nature Reserve, Glamorgan, Wales, UK. June.

Nick Upton / Red-tailed bumblebee (Bombus lapidarius) worker flying towards nest burrow in mature, stable sand dunes, Kenfig National Nature Reserve, Glamorgan, Wales, UK. June.

Nick Upton / Red-tailed bumblebee (Bombus lapidarius) worker with fully laden pollen baskets entering nest burrow in mature, stable sand dunes, Kenfig National Nature Reserve, Glamorgan, Wales, UK. June.

Nick Upton / Buff-tailed bumblebee (Bombus terrestris) nectaring on Hawthorn (Crataegus monogyna) blossom, Kenfig National Nature Reserve, Glamorgan, Wales, UK, May.

Nick Upton / Bumblebee (Bombus sp.) visiting a flowering, four metre tall Tree echium / Giant viper's bugloss (Echium pininana) in a garden, The Lizard Peninsula, Cornwall, UK. June.

Nick Upton / Buff-tailed bumble bee (Bombus terrestris) nectaring on Hound's tongue (Cynoglossum officinale) flowering on sand dunes, Merthyr Mawr National Nature Reserve, Glamorgan, Wales, UK. May.

Nick Upton / Bumblebee (Bombus sp.) visiting Viper's bugloss (Echium vulgare) flowers on sand dunes, Merthyr Mawr Warren National Nature Reserve, Glamorgan, Wales, UK. July.

Nick Upton / Female Vernal colletes / Spring mining bee (Colletes cunicularius) sunning itself on sand on a coastal sand dune, Merthyr Mawr National Nature Reserve, Glamorgan, Wales, UK. April.

Nick Upton / Female Vernal colletes / Spring mining bee (Colletes cunicularius) entering its nest burrow in a coastal sand dune, Merthyr Mawr National Nature Reserve, Glamorgan, Wales, UK. April.

Nick Upton / Eroded bank in a coastal sand dune with many active burrows of solitary bees, including the Vernal colletes / Spring mining bee (Colletes cunicularius), Merthyr Mawr National Nature Reserve, Glamorgan...

Nick Upton / Male Sandpit mining bee (Andrena barbilabris) sunning on a coastal sand dune, Merthyr Mawr National Nature Reserve, Glamorgan, Wales, UK. April.

Nick Upton / Female Silvery leafcutter bee (Megachile leachella) female approaching its nest burrow in coastal sand dunes, Kenfig National Nature Reserve, Glamorgan, Wales, UK, June.

Nick Upton / Female Silvery leafcutter bee (Megachile leachella) flying to its burrow in coastal dunes with circle of leaf it has cut to line its nest cells with, Dorset, UK. July.

Nick Upton / Female Silvery leafcutter bee (Megachile leachella) entering its burrow in coastal dunes with circle of leaf it has cut to line its nest cells with, Dorset, UK. July.

Nick Upton / Female Silvery leafcutter bee (Megachile leachella) returning to her burrow in coastal dunes with mass of pollen attached to the hairs under her abdomen to stick her brood cells with, Dorset, UK. July...

Nick Upton / Silvery leafcutter bee (Megachile leachella) excavating its burrow in coastal dunes, raking and kicking sand away with its legs, Dorset, UK. July.

Nick Upton / Female Silvery leafcutter bee (Megachile leachella) emerging from her burrow in coastal dunes, Dorset, UK. July.

Nick Upton / Hairy-saddled Colletes bee (Colletes fodiens) emerging from its nest burrow in a sandy bank, Merthyr Mawr National Nature Reserve, Glamorgan, Wales, UK, September.

Nick Upton / Hairy-saddled Colletes bee (Colletes fodiens) near its nest site in a sandy bank, Merthyr Mawr National Nature Reserve, Glamorgan, Wales, UK, September.

Nick Upton / Coast leafcutter bee (Megachile maritima) entering its burrow in coastal sand dunes, Dorset, UK, August.

Nick Upton / Silvery leafcutter bee (Megachile leachella) visiting a Sea bindweed (Calystegia soldanella) flower on coastal dunes, it's typical habitat, Dorset, UK, June.

Nick Upton / Mason bee / Leafcutter bee (Megachile / Chalicodoma sicula) nectaring on Sand viper's bugloss (Echium salbulicola) flowering on sandy coastal scrubland, Mondrago Natural Park, Majorca south coast, May...