Ingo Arndt / Grizzly Bear (Ursus arctos horribilis) front and rear tracks in mud, Lake Clark National Park, Alaska

Ole Jorgen Liodden / Polar Bear (Ursus maritimus) sitting on floe in vast thawing sea-ice landscape. Spitsbergen, Svalbard, August 2011. Second place, Mammals Category, GDT 2012 competion. Winner in ANIMALS IN THEIR ENVIR...

Gerrit Vyn / Human footprint and oiled beach from the BP Deepwater Horizon oil leak in the Gulf of Mexico. Baldwin County, Alabama. USA, June 2010.

Jeff Wilson / Camera crew searching for crevasse to film in, Gorner glacier, near Zermatt in the Swiss Alps, Switzerland. For BBC Planet Earth series, Mountains episode, 2006

Nick Garbutt / Grizzly Bears (Ursus arctos horribilis) footprint along the Atnarko River, Tweedsmuir Park, British Columbia, Canada, September 2009

Robin Chittenden / Wood pigeon (Columba palumbus) footprints in snow, Norfolk, England, March

Laurie Campbell / Wader footprints in sand, Aberlady Bay, Firth of Forth, Scotland, UK, December

Igor Shpilenok / Kamchatka Brown bear (Ursus arctos beringianus) footprint in mud, Koryaksky Zapovednik, Russia.

Sue Flood / Polar bear footprint in snow {Ursus maritimus} Churchill Canada

Ingo Arndt / Grizzly Bear (Ursus arctos horribilis) track in mud, Lake Clark National Park, Alaska

Mark Carwardine / Sumatran rhino (Dicerorhinus sumatrensis) plaster cast of footprint. Way Kambas National Park, Lampung Province, southern Sumatra, Indonesia Critically endangered / threatened species - fewer than 20...

Mark Carwardine / Blue whale (Balaenoptera musculus) fluke print or footprint, the distinctive impression left behind after a dive, Baja California, Sea of Cortez (Gulf of California), Mexico, Endangered species

Vladimir Medvedev / Footprint of a Siberian tiger (Panthera tigris altaica) on fresh snow. Lazovskiy zapovednik, Primorskiy krai, Far East Russia. November 2004

Donald M. Jones / Desert Cottontail (Sylvilagus audubonii) tracks in sand, Nevada

Jim Brandenburg / Coyote (Canis latrans) tracks and oak leaves in snow, North America

Nick Upton / 2020VISION / Volunteer Shen Yan Liow looks on as mammalogist Gill Brown matches up inky hedgehog footprints left in a footprint tunnel with identification chart during Arnos Vale Cemetery Bioblitz, Bristol, UK, Ma...

Nick Upton / 2020VISION / Volunteer Shen Yan Liow looks on as mammalogist Gill Brown matches up inky hedgehog footprints left in a footprint tunnel with identification chart during Arnos Vale Cemetery Bioblitz, Bristol, UK, Ma...

Nick Upton / 2020VISION / Volunteer Shen Yan Liow looks on as mammalogist Gill Brown refers to a mammal footprint identification chart during Arnos Vale Cemetery Bioblitz, Bristol, UK, May 2012 Model released.

Nick Upton / 2020VISION / Members of the public and volunteers gathered to inspect mammal skulls and footprint casts shown by mammalogist Gill Brown during Arnos Vale Cemetery Bioblitz, Bristol, UK, May 2012

Jim Brandenburg / Fox tracks on frozen lake, Superior National Forest, Minnesota

Paul Williams / Jaguar (Panthera onca) pug mark next to Apple snail (Pomacea canaliculata) Pantanal, Pocone, Brazil

Juan Carlos Munoz / Fossil of footprint of three toed prehistoric reptile, Iguanodon, Spain

Shibai Xiao / Leopard cat (Prionailurus bengalensis) footprint in sand near the valley, Linzhi, Tibet, China, October 2010.

Michael Quinton / Red Fox (Vulpes vulpes) smelling tracks in snow, Alaska

Roland Seitre / Footprint of African Elephant (Loxodonta africana) in sand. Tsavo National Park, Kenya, 2009.

Pete Oxford / Jaguar (Panthera onca) footprints and handprint, Yasuni National Park, Amazon, Ecuador

Peter Cairns / 2020VISION / Human footprint in mudflats, Morecambe Bay, Cumbria, England, UK, February

Eric Dragesco / Snow Leopard (Panthera uncia) footprint / pug mark in snow, with tape measure for scale, Altai Mountains, Mongolia,

Matthias Breiter / Grizzly Bear (Ursus arctos horribilis) front and back paw prints, Katmai National Park, Alaska

Klaus Echle / Wildcat (Felis silvestris) footprint in snow. Rhine Valley near Frieburg, Germany, January.

Eric Baccega / Female Polar Bear (Ursus maritimus) footprint. Wapusk National Park, Manitoba, Churchill, March.

Eric Baccega / Female Polar Bear (Ursus maritimus) footprint. Wapusk National Park, Manitoba, Churchill, March.

Frederique Olivier / Footprint of Giant Petrel (Macronectes giganteus) in sand, Macquarie Island, Southern Pacific, Australian Antarctica.

Matthias Breiter / Wolf (Canis lupus) tracks, Katmai National Park, Alaska

David Woodfall / Dinosaur footprint in Triassic rocks. UK, October 2010

Stephen Belcher / Komodo Dragon (Varanus komodoensis) tracks in sand, Komodo Island, Komodo National Park, Indonesia

Steven Kazlowski / Polar bear (Ursus maritimus) footprint left in the mud by a hind foot, Barter Island, 1002 area of the Arctic National Wildlife Refuge, Alaska

Pete Oxford / South American River Turtle (Podocnemis expansa) tracks, Orinoco River, Apure, Venezuela

Pete Oxford / South American River Turtle (Podocnemis expansa) tracks being followed by researcher, Orinoco River, Apure, Venezuela

Pete Oxford / Yellow-spotted Amazon River Turtle (Podocnemis unifilis) tracks from female that came to shore to lay eggs, Orinoco River, Apure, Venezuela

Pete Oxford / Yellow-spotted Amazon River Turtle (Podocnemis unifilis) tracks from female that came to shore to lay eggs, Orinoco River, Apure, Venezuela

Pete Oxford / South American River Turtle (Podocnemis expansa) tracks, Orinoco River, Apure, Venezuela

Pete Oxford / Jaguar footprints (Panthera onca) Cuiaba river, Pantanal. Mato Grosso do Sul Province. Brazil There is much size variation among these cats but in the Pantanal they are larger than those found in th...

Sergey Gorshkov / Wolverine (Gulo gulo) tracks in snow, Kamchatka, Russia

Eric Baccega / Footprint of Arctic wolf (Canis lupus arctos) Banks Island, North West Territories, Canada

Pete Oxford / Footpath through dunes of Erg Chebbi near the village of Merzouga, Morocco

Pete Oxford / Pacific Green Sea Turtle (Chelonia mydas agassizi) tracks made by female coming up on land to lay eggs, Espumilla Beach, Santiago Island, Galapagos Islands, Ecuador

Pete Oxford / Pacific Green Sea Turtle (Chelonia mydas agassizi) tracks made by female returning to sea, Espumilla Beach, Santiago Island, Galapagos Islands, Ecuador

Patricio Robles Gil / Front paw track of Kamchatka brown bear (Ursus arctos beringianus) in sand on the bank of Kamchatka River, Kamchatka, Russia

Ingo Arndt / Red Fox (Vulpes vulpes) tracks on frozen lake, Germany

Pete Oxford / Spanish Lynx (Lynx pardinus) tracks, Donana National Park, Huelva, Andalusia, Spain

Pete Oxford / Spanish Lynx (Lynx pardinus) track and human hand print, Donana National Park, Huelva, Andalusia, Spain

Ingo Arndt / Black-backed Jackal (Canis mesomelas) tracks, Cape Cross, Namibia

Warwick Sloss / Polar bear {Ursus maritimus} footprint in the snow, Churchill, Manitoba, Canada.

Donald M. Jones / Timber Wolf (Canis lupus) tracks in mud, western Montana

Donald M. Jones / Grizzly Bear (Ursus arctos horribilis) tracks in sand, southern Alaska

Donald M. Jones / Grizzly Bear (Ursus arctos horribilis) track, southern Alaska

Stephen Belcher / Sumatran Rhinoceros (Dicerorhinus sumatrensis) foot prints in the jungle, Way Kambas National Park, Sumatra, Indonesia

Andy Rouse / Making footprint casts during Tiger census, Ranthambore NP, Rajasthan, India (non-ex)

Steven David Miller / 130 million year old megalosaur dinosaur footprints can be seen at low tide at Gantheaume Point, Broome, Western Australia

Steven David Miller / 130 million year old megalosaur dinosaur footprints can be seen at low tide at Gantheaume Point, Broome, Western Australia

Steven David Miller / 130 million year old megalosaur dinosaur footprints can be seen at low tide at Gantheaume Point, Broome, Western Australia .

Sebastian Kennerknecht / Snowy Plover (Charadrius nivosus) breeding sign with footsteps, Scott Creek Beach, Santa Cruz, California

Donald M. Jones / Wild Turkey (Meleagris gallopavo) tracks in snow, western Montana

Juan Carlos Munoz / Pugmark of Siberian tiger {Panthera tigris altaica} in snow, captive, China

Jouan & Rius / Kangaroo footprint in sand dune, Mungo National Park, New South Wales, Australia

John Cancalosi / Fossil dinosaur footprint (Theropod, possibly Dilophosaurus sp) from Early Jurassic period, 200 million years, Navajo reservation near Tuba city Arizona, USA

John Cancalosi / Fossil dinosaur footprint (Theropod, possibly Dilophosaurus sp) from Early Jurassic period, 200 million years, Navajo reservation near Tuba city Arizona, USA

Steven Kazlowski / Polar bear (Ursus maritimus) footprint / track on Bernard Spit, a barrier island off the Arctic National Wildlife Refuge, Alaska

Steven Kazlowski / Polar bear (Ursus maritimus) cub footprint in snow, off the Arctic National Wildlife Refuge, Alaska

Michael & Patricia Fogden / Dinosaur tracks, Mount Etjo, Namibia

Fabio Liverani / Footprint of European brown bear {Ursus arctos} in the mud, Notranska, Slovenia

Fabio Liverani / Footprint of European brown bear {Ursus arctos} in the mud, Notranska, Slovenia

Yva Momatiuk & John Eastcott / King Penguin (Aptenodytes patagonicus) walking on steep and snowy alpine slope, Antarctic Bay, South Georgia Island

Yva Momatiuk & John Eastcott / King Penguin (Aptenodytes patagonicus) walking through snow, Antarctic Bay, South Georgia Island

Yva Momatiuk & John Eastcott / King Penguin (Aptenodytes patagonicus) trio on beach, St. Andrews Bay, South Georgia Island

Ingo Arndt / Grizzly Bear (Ursus arctos horribilis) tracks on tidal flats, Alaska

Ingo Arndt / Grizzly Bear (Ursus arctos horribilis) tracks on tidal flats, Alaska

Suzi Eszterhas / Grizzly Bear (Ursus arctos horribilis) tracks in mud, Katmai National Park, Alaska

Suzi Eszterhas / Grizzly Bear (Ursus arctos horribilis) tracks in mud, Katmai National Park, Alaska

Scott Leslie / Plover footprints on beach, Nova Scotia, Canada

Dietmar Nill / Two human footprints in wet sand, Baltic Sea coast of Poland

Anup Shah / Footprint of African lion {Panthera leo} in mud, Masai Mara GR, Kenya

Shattil & Rozinski / Grey wolf footprint {Canis lupus} beside man's hand for size comparison, Alaska, USA

Juergen Freund / 130 million year old Dinosaur footprints, Gantheaume Point, Broome, Western Australia

Matthias Breiter / Dinosaur prints, Gantheaume Point, Broome, Australia

Pete Oxford / Jaguar (Panthera onca) footprints, Cuiaba River, Brazil

Tim Fitzharris / Coyote tracks flowering shrubs on the dune fields in front of the Sangre de Cristo Mountains, Great Sand Dunes National Monument, Colorado

Pete Oxford / Human footprints in the sand. Cerro Brujo, San Cristóbal Island. GALAPAGOS

Michael Durham / Sumatran Tiger (Panthera tigris sumatrae) footprint along the Dato Ghani Trail in Endau-Rompin National Park, Malaysia

Flip Nicklin / Polar Bear (Ursus maritimus) track in snow, Canada

Michael Durham / Footprint in beach sand, Oregon coast

Tim Fitzharris / Dune vegetation and shorebird tracks in sand, Santa Rosa Island, Gulf Islands National Seashore, Florida

Pete Oxford / Ocelot {Felis pardalis} footprint in mud, Ecuador

Bernard Castelein / Tiger footprints (Panthera tigris) Kanha NP, Madhya Pradesh, India