Lucas Bustamante / Choco Toucan (Ramphastos brevis) on a tree with epiphytes, Mashpi, Pichincha, Ecuador

John Shaw / Chestnut-eared aracari (Pteroglossus castanotis) on cat-gut vine. Pantanal, Brazil, August.

David Pattyn / Plate-billed Mountain-Toucan (Andigena laminirostris), Bellavista private reserve, Mindo cloud forest, Pichincha, Ecuador, July

Daniel Brockner/ BIA / Crimson-rumped Toucanet (Aulacorhynchus haematopygus), Mindo, Ecuador

Paul Hobson / Keel-billed toucan (Ramphastos sulfuratus) perched on branch with beak open. Laguna del Lagarto, Alajuela, Costa Rica.

Thomas Marent / Keel-billed Toucan (Ramphastos sulfuratus), native to Central and South America

Thomas Marent / Red-headed Barbet (Eubucco bourcierii) male, Valle del Cauca, Colombia

Nick Garbutt / Toco toucan (Ramphastos toco) acrobatically perched , feeding in forest canopy. Northern Pantanal, Cuiaba River, Mato Grosso, Brazil.

Glenn Bartley/ BIA / Keel-billed Toucan (Ramphastos sulfuratus), Costa Rica

Glenn Bartley/ BIA / Black-mandibled Toucan (Ramphastos ambiguus) during rainfall, Costa Rica

Chien Lee / Plate-billed Mountain-Toucan (Andigena laminirostris), Mindo, Ecuador

Chien Lee / Plate-billed Mountain-Toucan (Andigena laminirostris), Mindo, Ecuador

Glenn Bartley/ BIA / Versicolored Barbet (Eubucco versicolor) male, Amazon, Peru

Glenn Bartley/ BIA / Collared Aracari (Pteroglossus torquatus) during rainfall, Costa Rica

Glenn Bartley/ BIA / Saffron Toucanet (Baillonius bailloni), Atlantic Rainforest, Brazil

Pete Oxford / Pale-mandibled Aracari (Pteroglossus erythropygius), Mashpi Rainforest Biodiversity Reserve, Pichincha, Ecuador

Pete Oxford / Pale-mandibled Aracari (Pteroglossus erythropygius), Mashpi Rainforest Biodiversity Reserve, Pichincha, Ecuador

Pete Oxford / Pale-mandibled Aracari (Pteroglossus erythropygius) trio, Mashpi Rainforest Biodiversity Reserve, Pichincha, Ecuador

Bence Mate / Collared aracari (Pteroglossus torquatus) squabbling with Montezuma oropendola (Psarocolius montezuma) Costa Rica.

Bence Mate / Collared aracari (Pteroglossus torquatus) two perched, possibly courtship feeding, Costa Rica.

Hermann Brehm / Toco Toucan (Ramphastos toco) , Pantanal, Brazil.

Wim van den Heever / Toco toucan (Ramphastos toco) in flight, Pantanal, Brazil.

Wim van den Heever / Toco toucan (Ramphastos toco) portrait, Pantanal Brazil.

Suzi Eszterhas / Keel-billed Toucan (Ramphastos sulfuratus), Costa Rica

Suzi Eszterhas / Keel-billed Toucan (Ramphastos sulfuratus), Costa Rica

Tui De Roy / Plate-billed Mountain-Toucan (Andigena laminirostris) at nest cavity, Bellavista Cloud Forest Reserve, Tandayapa Valley, Andes, Ecuador

Tui De Roy / Plate-billed Mountain-Toucan (Andigena laminirostris), Andes, Ecuador

Tui De Roy / Plate-billed Mountain-Toucan (Andigena laminirostris) at nest cavity, Bellavista Cloud Forest Reserve, Tandayapa Valley, Andes, Ecuador

Tui De Roy / Crimson-rumped Toucanet (Aulacorhynchus haematopygus), Angel Paz Reserve, Andes, Ecuador

Tui De Roy / Crimson-rumped Toucanet (Aulacorhynchus haematopygus), Bellavista Cloud Forest Reserve, Tandayapa Valley, Andes, Ecuador

Angelo Gandolfi / Toco Toucan (Ramphastos toco) beak open with tongue visible while feeding on mango, Brazil

Glenn Bartley / Many-banded Aracari (Pteroglossus pluricinctus), Ecuador

Glenn Bartley / Chestnut-mandibled Toucan (Ramphastos swainsonii), Costa Rica

Glenn Bartley / Emerald Toucanet (Aulacorhynchus prasinus), Costa Rica

Claudio Contreras / Keel billed toucan (Ramphastos sulfuratus) calling, captive.

Claudio Contreras / Keel billed toucan (Ramphastos sulfuratus), captive.

Bence Mate / Keel billed toucan (Ramphastos sulfuratus) in aggressive encounter with Montezuma Oropendola (Psarocolius montezuma) in rain, Laguna del Lagarto, Santa Rita, Costa Rica. Front cover bookplate of the H...

Nick Garbutt / Toco Toucans (Ramphastos toco) taking flight from the forest canopy. Banks of the Cuiaba River, northern Pantanal, Mato Grosso, Brazil.

Nick Garbutt / Toco Toucan (Ramphastos toco) feeding on fruit in forest canopy. Banks of the Cuiaba River, northern Pantanal, Mato Grosso, Brazil.

Suzi Eszterhas / Collared Aracari (Pteroglossus torquatus), northern Costa Rica

Murray Cooper / Plate-billed Mountain-Toucan (Andigena laminirostris) with orchids, Ecuador

Tui De Roy / Toucan Barbet (Semnornis ramphastinus) beside a bromeliad, Bellavista Cloud Forest Reserve, Ecuador

Tui De Roy / Crimson-rumped Toucanet (Aulacorhynchus haematopygus) eating fruit at feeding station, Bellavista Cloud Forest Reserve, Ecuador

Tim Fitzharris / Keel-billed Toucan (Ramphastos sulfuratus) pair, Costa Rica

Kevin Schafer / Red-breasted Toucan (Ramphastos dicolorus) calling, Itatiaia National Park, Brazil

Michael & Patricia Fogden / Keel-billed Toucan (Ramphastos sulfuratus) portrait, rainforest in Costa Rica

Michael & Patricia Fogden / Keel-billed Toucan (Ramphastos sulfuratus) perched in tree, Costa Rican rainforest

Michael & Patricia Fogden / Keel-billed Toucan (Ramphastos sulfuratus) perched in tree, Costa Rican rainforest

Michael & Patricia Fogden / Emerald Toucanet (Aulacorhynchus prasinus) eating Passion Fower (Passiflora sp) fruit at nest cavity, cloud forest, Costa Rica

Claus Meyer / Toco Toucan (Ramphastos toco) calling while perching on branch, Cerrado, Brazil

Flip de Nooyer / Black-necked Aracari (Pteroglossus aracari) a toucanet, perching on branch, Pantanal, Brazil

Gerry Ellis / Keel-billed Toucan (Ramphastos sulfuratus) portrait showing barbed tongue, native to Mexico and Central America

Claus Meyer / Red-breasted Toucan (Ramphastos dicolorus) close-up through vegetation, south Brazil

Norbert Wu / Toco Toucan (Ramphastos toco) bill is actually light and spongy, Brazil

Norbert Wu / Toco Toucan (Ramphastos toco) bill is actually light and spongy, Brazil

Mark Moffett / Emerald Toucanet (Aulacorhynchus prasinus), Monteverde Cloud Forest Reserve, Costa Rica

Staffan Widstrand / Wild Wonders of China / Great barbet (Psilopogon virens) Gaoligongshan, Yunnan, China

Staffan Widstrand / Wild Wonders of China / Blue-throated barbet (Psilopogon asiaticus) Tongbiguan Nature Reserve, Dehong, Yunnan, China

Staffan Widstrand / Wild Wonders of China / Blue-throated barbet (Psilopogon asiaticus) Tongbiguan Nature Reserve, Dehong, Yunnan, China

Gabriel Rojo / Toco toucan (Ramphastos toco) pair billing in tree. Pantanal forest, Mato Grosso, Brazil.

Lucas Bustamante / Many-banded aracari (Pteroglossus pluricinctus) Yasuni National Park, Orellana, Ecuador

John Shaw / Crimson-rumped toucanet (Aulacorhynchus haematopygus) perched, Ecuador, May.

Bernard Castelein / Chestnut-eared Aracari (Pteroglossus castanotis) Pampas del Yacuma Protected Area, Bolivia

David Pattyn / Plate-billed Mountain-Toucan (Andigena laminirostris) feeding on fruit, Bellavista private reserve, Mindo cloud forest, Pichincha, Ecuador, July

David Pattyn / Many-banded aracari (Pteroglossus pluricinctus) Cuyabeno wildlife reserve, Sucumbios, Amazon rainforest, Ecuador. July

David Pattyn / Many-banded aracari (Pteroglossus pluricinctus) Cuyabeno wildlife reserve, Sucumbios, Amazon rainforest, Ecuador. July

David Pattyn / Crimson rumped toucanet (Aulacorhynchus haematopygus) Mindo, Pichincha, Ecuador, July

Thomas Marent / Crimson-rumped Toucanet (Aulacorhynchus haematopygus), Valle del Cauca, Colombia

Thomas Marent / Crimson-rumped Toucanet (Aulacorhynchus haematopygus), Valle del Cauca, Colombia

Thomas Marent / Crimson-rumped Toucanet (Aulacorhynchus haematopygus), Valle del Cauca, Colombia

Thomas Marent / Crimson-rumped Toucanet (Aulacorhynchus haematopygus), Valle del Cauca, Colombia

Thomas Marent / Crimson-rumped Toucanet (Aulacorhynchus haematopygus) pair, Valle del Cauca, Colombia

Thomas Marent / Emerald Toucanet (Aulacorhynchus prasinus), Valle del Cauca, Colombia

Thomas Marent / Emerald Toucanet (Aulacorhynchus prasinus), Selva de Ventanas Natural Reserve, Colombia

Thomas Marent / Black-billed Mountain-Toucan (Andigena nigrirostris) calling, Selva de Ventanas Natural Reserve, Colombia

Thomas Marent / Black-billed Mountain-Toucan (Andigena nigrirostris), Selva de Ventanas Natural Reserve, Colombia

Thomas Marent / Black-billed Mountain-Toucan (Andigena nigrirostris) calling, Selva de Ventanas Natural Reserve, Colombia

Thomas Marent / Black-billed Mountain-Toucan (Andigena nigrirostris), Selva de Ventanas Natural Reserve, Colombia

Thomas Marent / Black-mandibled Toucan (Ramphastos ambiguus), Colombia

Thomas Marent / Channel-billed Toucan (Ramphastos vitellinus), Colombia

Thomas Marent / Channel-billed Toucan (Ramphastos vitellinus), Colombia

Thomas Marent / Channel-billed Toucan (Ramphastos vitellinus), Colombia

Thomas Marent / Keel-billed Toucan (Ramphastos sulfuratus), native to Central and South America

Thomas Marent / Keel-billed Toucan (Ramphastos sulfuratus), native to Central and South America

Thomas Marent / Red-headed Barbet (Eubucco bourcierii) male, Valle del Cauca, Colombia

Thomas Marent / Red-headed Barbet (Eubucco bourcierii) female, Valle del Cauca, Colombia

Thomas Marent / White-mantled Barbet (Capito hypoleucus), Rio Claro Nature Reserve, Colombia

Thomas Marent / White-mantled Barbet (Capito hypoleucus), Rio Claro Nature Reserve, Colombia

Philippe Clement / Blue-throated toucanet (Aulacorhynchus caeruleogularis) native to Costa Rica, Panama and Colombia, captive, digital composite

Glenn Bartley/ BIA / Keel-billed Toucan (Ramphastos sulfuratus), Costa Rica

Glenn Bartley/ BIA / Red-headed Barbet (Eubucco bourcierii) female, Andes, Colombia

Glenn Bartley/ BIA / Red-headed Barbet (Eubucco bourcierii) male, Andes, Colombia

Glenn Bartley/ BIA / Black-mandibled Toucan (Ramphastos ambiguus) during rainfall, Costa Rica

David Perpinan / Red breasted toucan (Ramphastos dicolorus) in cage, confiscated from illegal wildlife trade, Brazil.

Staffan Widstrand / Wild Wonders of China / Golden-throated barbet (Psilipogon franklinii) bird perched on a branch of a tree in Baihualing, Gaoligongshan, Yunnan, China