Pete Oxford / Spotted Scorpionfish (Scorpaena plumieri) camouflaged in rocks near King Angelfish (Holacanthus passer), Tower Island, Galapagos Islands, Ecuador

Claudio Contreras / Giant Manta Ray (Manta birostris) at cleaning station with Clarion Angelfish (Holacanthus clarionensis) San Benedicto Island, Revillagigedo Archipelago Biosphere Reserve (The Socorro Islands), Pacific...

Claudio Contreras / Queen Angelfish (Holacanthus ciliaris) among other coral fish. Cancun National Park, Caribbean Sea, Mexico.

Alex Mustard / Rock beauty angelfish (Holacanthus tricolor) pair at dusk performing their courtship dance with the larger male circling the female, Georgetown, Grand Cayman, Cayman Islands, British West Indies. Cari...

Jeff Rotman / Queen angelfish close-up of tail base {Holacanthus ciliaris} Bahamas, Caribbean

Flip Nicklin / Queen Angelfish (Holacanthus ciliaris) juvenile, Cocos Island, Costa Rica

Constantinos Petrinos / Queen angelfish at reef {Holacanthus ciliaris} Bonaire, Caribbean.

Doug Perrine / Queen angelfish {Holacanthus ciliaris} over brain coral, Cayman Islands, Caribbean Sea, Atlantic Ocean (Non-ex).

Pascal Kobeh / Queen angelfish or blue or golden angelfish (Holacanthus ciliaris) Cozumel Island, Yucatan peninsula, Mexico.

Magnus Lundgren / Wild Wonders of China / Semicircle angelfish (Pomacanthus semicirculatus) juvenile, Kenting National Park, Hengchun Peninsula of Pingtung County, Taiwan

David Fleetham / Lemonpeel angelfish (Centropyge flavissimus) Indonesia.

David Fleetham / Queen angelfish (Holacanthus ciliaris) on coral reef, Bonaire, Caribbean.

Linda Pitkin / Yellowbar angelfish (Pomacanthus maculosus) on coral reef, with Anthias (Pseudanthias squamipinnis) Shark Reef to Jolande, Ras Mohammed National Park, Egypt, Red Sea.

Linda Pitkin / Rock beauty fish (Holacanthus tricolor) sheltering against Blade Fire Coral (Millepora complanata) Bonaire, Leeward Antilles, Caribbean.

Linda Pitkin / Queen angelfish (Holacanthus ciliaris) Bonaire, Leeward Antilles, Caribbean.

Linda Pitkin / Queen angelfish (Holacanthus ciliaris) Bonaire, Leeward Antilles, Caribbean.

Alex Mustard / Pair of Rock beauty angelfish (Holacanthus tricolor) spawning at dusk. The larger male rubs against the flank of the female during the spawning rise. George Town, Grand Cayman, Cayman Islands, British...

Claudio Contreras / King angelfish (Holacanthus passer), Salsipuedes Island Protected Area, Gulf of California (Sea of Cortez), Mexico, July

Claudio Contreras / King angelfish (Holacanthus passer), San Pedro Martir Island Protected Area, Gulf of California (Sea of Cortez), Mexico, July

Georgette Douwma / Yellowbar angelfish (Pomacanthus maculosus). Red Sea, Egypt. January.

Georgette Douwma / Yellowbar angelfish (Pomacanthus maculosus), Red Sea, Egypt. January.

Georgette Douwma / Yellowbar angelfish (Pomacanthus maculosus), Red Sea, Egypt. January.

Tui De Roy / Marine Iguana (Amblyrhynchus cristatus) grazing on algae near King Angelfish (Holacanthus passer) school, Cape Douglas, Fernandina Island, Galapagos Islands, Ecuador

Nature Production / Portrait of Lemonpeel angelfish (Centropyge flavissima) front face, Chichijima, Ogasawara Islands UNESCO World Heritage Site, Japan

Claudio Contreras / Clarion angelfish (Holacanthus clarionensis) Socorro Island, Revillagigedo Archipelago Biosphere Reserve (Socorro Islands), Pacific Ocean, Western Mexico, March

Claudio Contreras / Flag Cabrilla (Epinephelus labriformis) and Clarion Angelfish (Holacanthus clarionensis), Clarion Angelfish IUCN Vulnerable, San Benedicto Island, Revillagigedo Archipelago Biosphere Reserve (Socorro...

Claudio Contreras / Carolines / Stareye Parrotfish (Calotomus carolinus), territorial dispute, Clarion Angelfish (Holacanthus clarionensis), Clarion Angelfish IUCN Vulnerable, San Benedicto Island, Revillagigedo Archipel...

Claudio Contreras / Clarion Angelfish (Holacanthus clarionensis) shoal, IUCN Vulnerable, Socorro Island, Revillagigedo Archipelago Biosphere Reserve / Archipielago de Revillagigedo UNESCO Natural World Heritage Site (Soc...

Claudio Contreras / Clarion angelfish (Holacanthus clarionensis) shoal, Socorro Island, Revillagigedo Archipelago Biosphere Reserve / Archipielago de Revillagigedo UNESCO Natural World Heritage Site (Socorro Islands), P...

Claudio Contreras / Clarion Angelfish (Holacanthus clarionensis) and coral, IUCN Vulnerable, Socorro Island, Revillagigedo Archipelago Biosphere Reserve / Archipielago de Revillagigedo UNESCO Natural World Heritage Site...

Claudio Contreras / Clarion Angelfish (Holacanthus clarionensis) group of three, IUCN Vulnerable, Socorro Island, Revillagigedo Archipelago Biosphere Reserve / Archipielago de Revillagigedo UNESCO Natural World Heritage...

Claudio Contreras / Clarion Angelfish (Holacanthus clarionensis), IUCN Vulnerable, Socorro Island, Revillagigedo Archipelago Biosphere Reserve / Archipielago de Revillagigedo UNESCO Natural World Heritage Site (Socorro I...

Alex Mustard / Majestic / Blue girdled angelfish (Pomacanthus navarchus) swims through hard coral. Kri Island, Raja Ampat, West Papua, Indonesia.

Alex Mustard / Pattern detail of a Majestic angelfish (Pomacanthus navarchus). Fiabacet Islands, Misool, Raja Ampat, West Papua, Indonesia. Ceram Sea.

Alex Mustard / Close up of the scale details of a Yellow-mask angelfish (Pomacanthus xanthometopon). Baa Atoll, Maldives. Indian Ocean

Georgette Douwma / Meyer's butterflyfish (Chaetodon meyeri) Andaman Sea, Thailand.

Georgette Douwma / Yellowbar angelfish (Pomacanthus maculosus) Egypt, Red Sea.

Georgette Douwma / Yellowbar angelfish (Pomacanthus maculosus) swimming over rocky reef, Egypt, Red Sea.

Georgette Douwma / Yellowbar angelfish (Pomacanthus maculosus) swimming over rocky reef, Egypt, Red Sea.

Georgette Douwma / Yellowbar angelfish (Pomacanthus maculosus) swimming over rocky reef, Egypt, Red Sea.

Georgette Douwma / Semicircular angelfish (Pomacanthus semicirculatus) swimming over coral reef. Komodo National Park, Indonesia. Indo-Pacific.

Georgette Douwma / Stoplight parrotfish (Sparisoma viride), terminal phase, grazing on coral rock, shadowed by a Trumpetfish (Aulostomus maculatus) next to a Rocky beauty (Holacanthus tricolor) Bonaire, Netherlands Ant...

Georgette Douwma / Queen angelfish (Holocanthus ciliaris) profile Bonaire, Netherlands Antilles, Caribbean, Atlantic Ocean.

Georgette Douwma / Queen angelfish (Holocanthus ciliaris) feeding on sponge. Bonaire, Netherlands Antilles, Caribbean, Atlantic Ocean.

Georgette Douwma / Queen angelfish (Holocanthus ciliaris) profile, Bonaire, Netherlands Antilles, Caribbean, Atlantic Ocean.

Georgette Douwma / Queen angelfish (Holocanthus ciliaris) feeding on sponge. Bonaire, Netherlands Antilles, Caribbean, Atlantic Ocean.

Georgette Douwma / Rocky beauty (Holacanthus tricolor) swimming near corals, Bonaire, Netherlands Antilles, Caribbean, Atlantic Ocean.

Georgette Douwma / Rocky beauty (Holacanthus tricolor) profile Bonaire, Netherlands Antilles, Caribbean, Atlantic Ocean.

Georgette Douwma / Queen angelfish (Holocanthus ciliaris) profile, Bonaire, Netherlands Antilles, Caribbean, Atlantic Ocean.

Georgette Douwma / Queen angelfish (Holocanthus ciliaris) with mouth open, Bonaire, Netherlands Antilles, Caribbean, Atlantic Ocean.

Georgette Douwma / Queen angelfish (Holocanthus ciliaris) feeding on sponge, whilst a Dusky damselfish (Stegastes adustus), a very territorial fish, tries to chase it away. Bonaire, Netherlands Antilles, Caribbean, Atl...

Pete Oxford / Regal angelfish (Pygoplites diacanthus) Rainbow Reef, Fiji, South Pacific.

Pascal Kobeh / Arabian angelfish (Pomacanthus maculosus) on reef, Sudan. Red Sea.

Pascal Kobeh / Giant manta ray (manta birostris) with two Remoras (Remora remora) attached its body and surrounded with Black jacks / trevally (Caranx lugubris) and a Clarion angelfish (Holacanthus clarionensis) Re...

Pascal Kobeh / Giant manta ray (manta birostris) with two Remoras (Remora remora) attached its body and surrounded with Black jacks / trevally (Caranx lugubris) and a Clarion angelfish (Holacanthus clarionensis) Re...

Pascal Kobeh / Diver filming a Giant manta ray (manta birostris) with two Remoras (Remora remora) attached its body and surrounded with Black jacks / trevally (Caranx lugubris) and a Clarion angelfish (Holacanthus c...

Pascal Kobeh / Divers behind a Giant manta ray (manta birostris) with two Remoras (Remora remora) attached its body and surrounded with Black jacks / trevally (Caranx lugubris) and a Clarion angelfish (Holacanthus c...

Pascal Kobeh / Giant manta ray (manta birostris) swimming in open water surrounded with black jacks / trevally (Caranx lugubris) and a Clarion angelfish (Holacanthus clarionensis) which usually cleans it, Revillagig...

Pascal Kobeh / Giant manta ray (manta birostris) swimming in open water surrounded with Black jacks / trevally (Caranx lugubris) and a Clarion angelfish (Holacanthus clarionensis) Revillagigedo islands, Mexico. Paci...

Pascal Kobeh / Giant manta ray (manta birostris) swimming in open water surrounded with Black jacks / trevally (Caranx lugubris) and a Clarion angelfish (Holacanthus clarionensis) Revillagigedo islands, Mexico. Paci...

Pascal Kobeh / Giant manta ray (manta birostris) swimming in open water surrounded with Black jacks / trevally (Caranx lugubris) and a Clarion angelfish (Holacanthus clarionensis) Revillagigedo islands, Mexico. Paci...

Pascal Kobeh / Giant manta ray (manta birostris) swimming in open water surrounded with Black jacks / trevally (Caranx lugubris) and a Clarion angelfish (Holacanthus clarionensis) Revillagigedo islands, Mexico. Paci...

Pascal Kobeh / Half-circled angelfish (Pomacanthus semicirculatus) Madagascar. Indian Ocean.

Georgette Douwma / Regal angelfish (Pygoplites diacanthus) swimming over coral reef. Egypt, Red Sea.

Georgette Douwma / Yellowbar angelfish (Pomacanthus maculosus) swimming over coral reef. Egypt, Red Sea.

Georgette Douwma / Yellowbar angelfish (Pomacanthus maculosus) swimming over coral reef. Egypt, Red Sea.

Georgette Douwma / Yellowbar angelfish (Pomacanthus maculosus) Egypt, Red Sea.

Jeff Rotman / Short spined porcupine fish (Diodon holacanthus) in Lembeh Strait, Celebes Sea, Sulawesi, Indonesia.

MYN / Paul Marcellini / Blue angelfish (Holacanthus isabelita) juvenile, Biscayne National Park, Florida, USA, May. meetyourneighbours.net project

MYN / Paul Marcellini / Blue angelfish (Holacanthus isabelita) juvenile, Biscayne National Park, Florida, USA, May. meetyourneighbours.net project

Claudio Contreras / Queen Angelfish (Holacanthus ciliaris). Cancun National Park, Caribbean Sea, Mexico.

Claudio Contreras / Rock Beauty (Holacanthus tricolor). Cancun National Park, Caribbean Sea, Mexico.

Claudio Contreras / Rock Beauty (Holacanthus tricolor). Cancun National Park, Caribbean Sea, Mexico.

Claudio Contreras / Blue Angelfish (Holacanthus bermudensis) juvenile, Cancun National Park, Caribbean Sea, Mexico.

Alex Mustard / Rock beauty angelfish (Holacanthus tricolor) swimming through a small arch on a colourful coral reef, East End, Grand Cayman, Cayman Islands, British West Indies, Caribbean Sea.

Alex Mustard / Rock beauty (Holacanthus tricolor) pair spawning, with the male rubbing against the flank of the female during the spawning rise, Georgetown, Grand Cayman, Cayman Islands, British West Indies, Caribbe...

Alex Mustard / Queen angelfish (Holacanthus ciliaris) portrait, swimming along the edge of a coral reef at dusk, Seven Mile Beach, Grand Cayman, Cayman Islands, British West Indies, Caribbean Sea.

Pete Oxford / Queen Angelfish (Holacanthus ciliaris) feeding on coral reef, Bonaire, Netherlands Antilles, Caribbean

Norbert Wu / Whitecheek Surgeonfish (Acanthurus nigricans) in school of Clarion Angelfish (Holacanthus clarionensis), Revillagigedos Islands, Mexico

Norbert Wu / Clarion Angelfish (Holacanthus clarionensis) school, Revillagigedos Islands, Mexico

Norbert Wu / Clarion Angelfish (Holacanthus clarionensis) group following Manta Ray (Manta birostris) to pick up food bits, Revillagigedos Islands, Mexico

Claudio Contreras / Blue angel fish (Holacanthus bermudensis) on coral reef, Cancun National Park, Caribbean Sea, Mexico, July

Fred Bavendam / Clarion Angelfish (Holacanthus clarionensis) juvenile, Socorro Island, Mexico

Fred Bavendam / King Angelfish (Holacanthus passer), Galapagos Islands, Ecuador

Fred Bavendam / Gulf Star (Oreaster occidentalis) pair and King Angelfish (Holacanthus passer) school, Galapagos Islands, Ecuador

Doug Perrine / King angelfish {Holacanthus passer} Galapagos

Doug Perrine / Natural hybrid cross between Blue angelfish {Holacanthus bermudensis} and Queen angelfish {Holacanthus ciliaris} Texas, Gulf of Mexico, USA

GEORGETTE DOUWMA / Rock beauty (Holacanthus tricolor) Bonaire, Netherlands Antilles, Caribbean

Pete Oxford / Shoal of King or White-banded angelfish (Holacanthus passer) Española / Hood Island, Galapagos

Dan Burton / Blue angelfish (Holacanthus bermudensis) Florida, USA

Pete Oxford / King Angelfish (Holacanthus passer) and Yellowtail Surgeonfish (Prionurus laticlavius) mixed school over reef, Gardner Bay, Espanola Island, Galapagos Islands, Ecuador

Peter Scoones / Queen Angelfish (Holocanthus ciliaris) on reef, Bonaire, Netherlands Antilles, Caribbean. photographed during making of BBC Planet Earth series 2005

Peter Scoones / Queen Angelfish (Holacanthus ciliaris) Bonaire, Netherlands Antilles, Caribbean photographed during making of BBC Planet Earth series 2005

Peter Scoones / Queen Angelfish (Holacanthus ciliaris) Bonaire, Netherlands Antilles, Caribbean photographed during making of BBC Planet Earth series 2005

Doug Perrine / Queen angelfish {Holacanthus ciliaris} Grand Bahama, Bahamas