Ashley Cooper / Hangdang Steel works, with billow of vapor, Hangdang, Northern China. March 2009.

Anup Shah / Western chimpanzees female 'Jire' aged 52 and her daughter 'Joya' aged 6 years and son 'Jeje' aged 13 years feeding on palm oil fruits stored by villagers, Bossou Forest, Mont Nimba, Guinea. December...

Christophe Courteau / Wild chimpanzees (Pan troglodytes schweinfurthii) in a fig tree, Kibale National Park, Uganda

Cheryl-Samantha Owen / A ranger looks for Chimpanzees in Semliki National Park's forest, Uganda, East Africa. No release available.

Suzi Eszterhas / Rescued chimpanzees (Pan troglodytes) playing with Bruce Ainebyona (Caretaker) Ngamba Island Chimpanzee Sanctuary, Uganda, Africa. Captive, June 2009. model released, *Digitally removed fence post in...

Anup Shah / Eastern common chimpanzee {Pan troglodytes schweinfurtheii} lying on ground, Mahale NP, Tanzania

Anup Shah / Chimpanzee in trees, 'Patti', Gombe NP, Tanzania 2002 {Pan troglodytes schweinfurtheii}

Anup Shah / Chimpanzee with young feeding on bark, Gombe NP, Tanzania 2003 'Tanga' + 'Tom'

Anup Shah / Young Chimpanzees + mother fishing for termites, Gombe NP, Tanzania 2003 'Gremlin'

Anup Shah / Chimpanzee swinging through trees, Gombe NP, Tanzania 2003 'Fanni'

Anup Shah / Chimpanzee + young in tree 'Gremlin' + 'G', Gombe NP, Tanzania 2003

Anup Shah / Male Chimpanzee yawning, Gombe NP, Tanzania 2003 'Tubbe' {Pan troglodytes schweinfurtheii}

Anup Shah / Male Chimpanzee + young fishing for termites, Gombe NP, Tanzania 2003 'Gremlin'

Anup Shah / Young Chimpanzee chewing vine, 'Tom', Gombe NP, Tanzania 2002

Karl Ammann / Young Chimpanzees using grass stems to extract termites from mound {Pan troglodytes} in sanctuary, Kenya

Anup Shah / Alpha male Chimpanzee 'Frodo', 26-years-old, feeding on leaves {Pan troglodytes schweinfurtheii} Kasekela community, Gombe NP, Tanzania. 2002. Chimps have learnt that certain leaves can be used to tre...

Anup Shah / Male Chimpanzee sitting on tree buttress, Gombe NP Tanzania. Apollo

Karl Ammann / Young Chimpanzees 'kissing' {Pan troglodytes} Chimfunshi sanctuary, Zambia

Jabruson / Young chimpanzee {Pan troglodytes schweinfurthii} Virunga NP, Democratic Republic of Congo.

Anup Shah / Chimpanzees feeding, Gombe NP, Tanzania. Fifi and Ferdinano

Martha Holmes / Chimpanzee {Pan troglodytes} hand on bar of cage.

Anup Shah / Juvenile Chimpanzee Flirt in tree {Pan troglodytes schweinfurthii} Gombe NP Tanzania

Anup Shah / Juvenile Chimpanzee 'Flirt' in tree {Pan troglodytes schweinfurthii} Gombe NP Tanzania

Jabruson / Chimpanzee looking at its reflection {Pan troglodytes} Ituri Forest, N.E.Zaire, Africa (chimp chimps chimpanzees)

Anup Shah / Female Chimpanzee 'Fanni' 20-years-old, resting with sons Fundi, 17-months-old, and Fudge, 5-years-old {Pan troglodytes schweinfurtheii} Gombe NP, Tanzania. 2002

Anup Shah / Chimpanzees gather round alpha male 'Frodo' in tree feeding on dead Bushbuck fawn {Pan troglodytes schweinfurtheii} Gombe NP, Tanzania. 2002

Anup Shah / Female Chimpanzee 'Gremlin' 31-years-old, carrying twin daughters Golden and Glitter, 3-years-3-monhts-old, jockey riding {Pan troglodytes schweinfurtheii} Gombe NP, Tanzania. 2002

Anup Shah / Young male Chimpanzee 'Gold' 3-years-3-months-old, watches mother 'Gremlin' use stick as tool to catch termites {Pan troglodytes schweinfurtheii} Gombe NP, Tanzania. 2002

Anup Shah / Chimpanzees mutual grooming {Pan troglodytes schweinfurtheii} Gombe NP, Tanzania. 2002

Anup Shah / Male Chimpanzee 'Paxo' 24-years-old, {Pan troglodytes schweinfurtheii} Gombe NP, Tanzania. 2002

Anup Shah / Young Chimpanzee hanging upside down in tree {Pan troglodytes schweinfurtheii} Gombe NP, Tanzania. 2002

Anup Shah / Chimpanzees gather round alpha male 'Frodo' in tree with dead Bushbuck fawn {Pan troglodytes schweinfurtheii} Gombe NP, Tanzania. 2002

Anup Shah / Chimpanzees mutual grooming {Pan troglodytes schweinfurtheii} Gombe NP, Tanzania. 2002

Anup Shah / Female Chimpanzee 'Fanni' with offspring 'Fundi' (male, 17-months-old) and Fudge (male, 4-years-10-months-old) {Pan troglodytes schweinfurtheii}, Kasekela community, Gombe NP, Tanzania. 2002

Anup Shah / Alpha male 'Frodo' displays 'keep away' message to begging chimps wanting share of Bushbuck fawn prey {Pan troglodytes schweinfurtheii}, Kasekela community, Gombe NP, Tanzania. 2002

Anup Shah / Male Chimpanzee 'Sheldon' knuckle walking carrying dead Red colobus monkey in jaws {Pan troglodytes schweinfurtheii}, 19-years-old, high ranking male of Kasekela community, Gombe NP, Tanzania. 2002

Anup Shah / Male Chimpanzee resting on rock 'Pax' {Pan troglodytes schweinfurtheii}, 25-years-old, lowest ranking male of Kasekela community, Gombe NP, Tanzania. 2002

Anup Shah / Alpha male Chimpanzee 'Frodo', 26-years-old {Pan troglodytes schweinfurtheii} Kasekela community, Gombe NP, Tanzania. 2002

Anup Shah / Female Chimpanzee 'Patti' (39-years-old) carries jockey-riding offspring 'Tarzan' on her back {Pan troglodytes schweinfurtheii}, Kasekela community, Gombe NP, Tanzania. 2002

Anup Shah / Female Chimpanzee 'Gremlin' demonstrates to offspring how to use a stem as a tool to remove termites from a termite mound {Pan troglodytes schweinfurtheii} Kasekela community, Gombe NP, Tanzania. 2002

Anup Shah / Fundi plays beside his resting mother 'Fanni' {Pan troglodytes schweinfurtheii}, Kasekela community, Gombe NP, Tanzania. 2002

Anup Shah / Two-year-old male 'Fundi' holds on to mother 'Fanni' who is calling in answer to group call {Pan troglodytes schweinfurtheii}, Kasekela community, Gombe NP, Tanzania. 2002 *Not available for calendar...

Jabruson / Young Chimpanzee playing with park warden {Pan troglodytes schweinfurthii} Virunga NP, Dem Rep of Congo

Jabruson / Male Chimpanzee (Opi) {Pan troglodytes schweinfurthii} Virunga NP, Democratic Republic of Congo.

Fiona Rogers / Western chimpanzee, male 'Tua' aged 53 years curious about a 'head support' used by villagers to carry heavy items made from palm leaves (Pan troglodytes verus). Bossou Forest, Mont Nimba, Guinea. Jan...

Ann & Steve Toon / Chimpanzees (Pan troglodytes) from West and Central Africa that were orphaned or abused find sanctuary in the Sweetwaters Chimpanzee Sanctuary, Ol Pejeta conservancy, Laikipia, Kenya, September 2012

Anup Shah / Western chimpanzees female 'Jire' aged 52 and her daughter 'Joya' aged 6 years feeding on palm oil fruits stored by villagers, Bossou Forest, Mont Nimba, Guinea. December 2010.

Anup Shah / Western chimpanzees female 'Jire' aged 52 and her daughter 'Joya' aged 6 years and son 'Jeje' aged 13 years feeding on palm oil fruits stored by villagers, Bossou Forest, Mont Nimba, Guinea. December...

Anup Shah / Western chimpanzees using leaves as sponge tools to drink from a hole in a tree, Bossou Forest, Mont Nimba, Guinea. January 2011.

Anup Shah / Western chimpanzee (Pan troglodytes verus) juvenile female 'Joya' aged 6 years throwing stones at other chimpanzees, Bossou Forest, Mont Nimba, Guinea. January 2011.

Anup Shah / Western chimpanzee (Pan troglodytes verus) male 'Foaf' aged 30 years making 'Pant hoot' vocalisations to make contact with other chimpanzees, Bossou Forest, Mont Nimba, Guinea. December 2010.

Anup Shah / Western chimpanzee (Pan troglodytes verus) juvenile female 'Joya' aged 6 years throwing rocks at other chimpanzees, Bossou Forest, Mont Nimba, Guinea. January 2011

Christophe Courteau / Fruit of the Fig tree (Ficus mucoso) fed on by Chimpanzees in the Kibale Rain Forest, Uganda

Christophe Courteau / Wild chimpanzees (Pan troglodytes schweinfurthii) grooming. On the right, female in oestrus with swollen genitalia, Kibale National Park, Uganda

Christophe Courteau / Kyambura Gorge, habitat of wild chimpanzees. Queen Elizabeth National Park, Uganda

Christophe Courteau / Two wild chimpanzees (Pan troglodytes schweinfurthii) sitting in a tree. The alfa / dominant male "Mobutu" is eating meat, the remains of a Black & White Collobus hunted by the whole group. Kibale Nat...

Christophe Courteau / *Person holding a sliced Star Apple / fruit of fig tree (Chrysophyllum albidum), a popular fruit in much of Africa, and a favourite of wild chimpanzees. Kibale Rainforest, Kibale National Park, Uganda

Christophe Courteau / Person holding a sliced Star Apple / fruit of fig tree (Chrysophyllum albidum), a popular fruit in much of Africa, and a favourite of wild chimpanzees. Kibale Rainforest, Kibale National Park, Uganda

Christophe Courteau / Star Apples / fruits of fig tree (Chrysophyllum albidum) amongst leaf litter on the forest floor, part-eaten by wild chimpanzees (Pan troglodytes schweinfurthii) in the Kibale Rainforest, Kibale Natio...

Christophe Courteau / Star Apples / fruits of fig tree (Chrysophyllum albidum) amongst leaf litter on the forest floor, part-eaten by wild chimpanzees (Pan troglodytes schweinfurthii) in the Kibale Rainforest, Kibale Natio...

Konrad Wothe / Chimpanzees (Pan troglodytes) old males, Mahale Mountains National Park, Tanzania, East Africa

Konrad Wothe / Chimpanzees (Pan troglodytes) aggressive males fight against their alpha male Pim, Mahale Mountains National Park, Tanzania, East Africa

Konrad Wothe / Chimpanzees (Pan troglodytes) aggressive males fight against their alpha male Pim, Mahale Mountains National Park, Tanzania, East Africa

Konrad Wothe / Chimpanzees (Pan troglodytes) aggressive males fight against their alpha male Pim, Mahale Mountains National Park, Tanzania, East Africa

Konrad Wothe / Chimpanzees (Pan troglodytes) aggressive males fight against their alpha male Pim, Mahale Mountains National Park, Tanzania, East Africa

Konrad Wothe / Chimpanzees (Pan troglodytes) aggressive males screaming, fight against their alpha male Pim, Mahale Mountains National Park, Tanzania, East Africa

Konrad Wothe / Chimpanzees (Pan troglodytes) aggressive males screaming, fight against their alpha male Pim, Mahale Mountains National Park, Tanzania, East Africa

Konrad Wothe / Male Chimpanzees fighting (Pan troglodytes) heavily injured alpha male 'Pim', Mahale Mountains National Park, Tanzania, East Africa

Konrad Wothe / Male Chimpanzees fighting, (Pan troglodytes) Mahale Mountains National Park, Tanzania, East Africa

Konrad Wothe / Chimpanzees (Pan troglodytes) aggressive males screaming, Mahale Mountains National Park, Tanzania, East Africa

Konrad Wothe / Chimpanzees (Pan troglodytes) aggressive males screaming, Mahale Mountains National Park, Tanzania, East Africa

Konrad Wothe / Chimpanzees (Pan troglodytes) females with baby, Mahale Mountains National Park, Tanzania, East Africa

Konrad Wothe / Chimpanzees (Pan troglodytes) mother grooming baby, Mahale Mountains National Park, Tanzania, East Africa

Konrad Wothe / Chimpanzees (Pan troglodytes) females grooming with baby, Mahale Mountains National Park, Tanzania, East Africa

Konrad Wothe / Chimpanzees (Pan troglodytes) females grooming with baby, Mahale Mountains National Park, Tanzania, East Africa

Konrad Wothe / Chimpanzees (Pan troglodytes) females grooming with baby, Mahale Mountains National Park, Tanzania, East Africa

Konrad Wothe / Chimpanzees (Pan troglodytes) mother with baby, Mahale Mountains National Park, Tanzania, East Africa

Konrad Wothe / Chimpanzees (Pan troglodytes) climbing, female with baby, Mahale Mountains National Park, Tanzania, East Africa

Nick Turner / Kyambura Gorge, habitat of wild chimpanzees. Queen Elizabeth National Park, Uganda, July 2010.

Simon Colmer / (Vernonia amygdalina) commonly called bitter leaf because of its bitter taste. In the wild, chimpanzees have been observed ingesting leaves to counter parasitic infections. Medicinal plant: used for w...

Florian Möllers / Male Chimpanzees (Pan troglodytes schweinfurthii) "Maani" (front) and "Kato" approaching in impressive posture, zoom-effect. Budongo Forest Reserve, Masindi, Uganda, Africa. December

Florian Möllers / Member of the Snare Removal Unit at Budongo Conservation Field Station removes snare set for duikers and bush pig that could seriously harm or mutilate chimpanzees which might accidentially get caug...

Florian Möllers / PhD student Zarin Machanda (Harvard University) and senior field assistant Geresomu Muhumuza tasting bitter food items of chimpanzees (Pan troglodytes schweinfurthii). Budongo Forest Reserve, Masindi,...

Florian Möllers / Male Chimpanzees (Pan troglodytes schweinfurthii) "Kato" (13 years, on left) and "Gashom" (18 years) sitting next to each other on a log in the rainforest. Budongo Forest Reserve, Masindi, Uganda, Afr...

Florian Möllers / Simon Townsend, PhD student (University of St. Andrews) and field assistant Monday Gideon, Zarin Machanda, PhD student (Harvard University) and Senior field assistant Geresomu Muhumuza observing chimp...

Florian Möllers / Field assistant Monday Gideon and PhD student Simon Townsend (University of St. Andrews) observing Chimpanzees (Pan troglodytes schweinfurthii) grooming on rainforest road. Budongo Forest Reserve, Mas...

Florian Möllers / Group of Chimpanzees (Pan troglodytes schweinfurthii) social grooming on forest floor. Budongo Forest Reserve, Masindi, Uganda, Africa. December

Florian Möllers / PhD student Simon Townsend (University of St. Andrews) and field assistant Monday Gideon observing Chimpanzees (Pan troglodytes schweinfurthii). Budongo Forest Reserve, Masindi, Uganda, Africa. Decemb...

Florian Möllers / PhD student Zarin Machanda (Harvard University) and senior field assistant Geresomu Muhumuza recording behavioural data from chimpanzees (Pan troglodytes schweinfurthii). Budongo Forest Reserve, Masin...

Florian Möllers / Senior field assistant Geresomu Muhumuza recording indiviudal behavioural data of chimpanzees (Pan troglodytes schweinfurthii). Budongo Forest Reserve, Masindi, Uganda, Africa. December 2006

Florian Möllers / Group of Chimpanzees (Pan troglodytes schweinfurthii) social grooming on forest floor. Budongo Forest Reserve, Masindi, Uganda, Africa. December

Florian Möllers / Simon Townsend, PhD student (University of St. Andrews) and field assistant Monday Gideon logging data of voice recordings from female chimpanzees (Pan troglodytes schweinfurthii). Budongo Forest Rese...

Florian Möllers / PhD student Zarin Machanda from Harvard University observing chimpanzees (Pan troglodytes schweinfurthii) together with field assistants. Budongo Forest Reserve, Masindi, Uganda, Africa. December 2006

Florian Möllers / Male Chimpanzees (Pan troglodytes schweinfurthii) "Zefa" (on left) and "Tinka" (45+ years). Budongo Forest Reserve, Masindi, Uganda, Africa. December

Florian Möllers / Moses Lemi of the Snare Removal Unit at Budongo Conservation Field Station remove snares that could seriously harm or mutilate chimpanzees which might accidentially get caught in them. Budongo Forest...

Florian Möllers / Figs around a large fig tree provide an important food source for Chimpanzees (Pan troglodytes schweinfurthii). Budongo Forest Reserve, Masindi, Uganda, Africa. December

Mark Carwardine / Keeper with rescued (orphaned) chimpanzees (Pan troglodytes) Ngamba Island Chimpanzee Sanctuary, Uganda, April 2007