Phil Savoie / Painted nomad bee (Nomada fucata) male bee with characteristic green eyes, sleeping with jaws locked around grass stem, Monmouthshire, Wales, UK. June

Loic Poidevin / Grizzly bear (Ursus arctos horribilis) mother watching cub catching salmon, Katmai National Park, Alaska, USA, August

Loic Poidevin / Grizzly bear (Ursus arctos horribilis) cubs with salmon, Katmai National Park, Alaska, August. Small reproduction only.

Loic Poidevin / Grizzly bear (Ursus arctos horribilis) cubs with salmon, Katmai National Park, Alaska, August. Small reproduction only.

John MacPherson / Reindeer (Rangifer tarandus) fighting, Cairngorms National Park, Scotland, UK, March.

Staffan Widstrand / Two Svalbard reindeer (Rangifer tarandus platyrhynchus) fighting; antlers shedding velvet; Spitsbergen, Svalbard, Norway, September

Denis-Huot / Two male Thomson's gazelles (Eudorcas thomsonii) fighting, Masai-Mara game reserve, Kenya.

Michel Roggo / Brown bear (Ursus arctos beringianus) fishing for Sockeye salmon in the Ozernaya River, Kamtchatka, Russia, August

Eric Medard / Ibex (Capra ibex) males fighting, Jura, Switzerland, captive

Alex Mustard,Alex Mustard / A female (in front) and male Sockeye salmon (Oncorhynchus nerka) over eggs in their spawning river. Adams River, British Columbia, Canada, October.

Alex Mustard / A split level photo of group of Sockeye salmon (Oncorhynchus nerka) fighting their way upstream as they migrate back to the river of their birth to spawn, trees showing autumnal colours. Adams River,...

Jack Dykinga / Rain spattered inter-locking palm leaves create a design of pattern and colour in the tropical rainforest. Osa Peninsula, Costa Rica