Anup Shah / Western lowland gorilla (Gorilla gorilla gorilla) sub-adult female 'Mosoko' aged 8 years crossing a river, Bai Hokou, Dzanga Sangha Special Dense Forest Reserve, Central African Republic

Nick Garbutt / Aggregation / bloom / swarm of Moon Jellyfish (Aurelia aurita) in the clear waters of Loch Na Keal, off the Isle of Mull, north west Scotland, UK, June, Photographed through the surface.

Bence Mate / Leaf cutter ant (Atta sp) female worker carrying leaf to nest, with a smaller 'minor' ant riding. The minor's job is to protect her sister from parasitic flies. Costa Rica, December. Highly Commended,...

Rolf Nussbaumer / Dung Beetle (Scarabaeinae) adult rolling dung ball. Laredo, Webb County, South Texas, USA Not available for ringtone/wallpaper use.

Gavin Hellier / New Pudong skyline, looking across the Huangpu River from the Bund, Shanghai, China 2010. No release available.

Gavin Hellier / In the famous souks of Marrakech, Morocco, 2011. No release available.

Gavin Hellier / Tuk Tuk or auto rickshaw in motion at night, Bangkok, Thailand, 2010. Model released.

Dale Sutton / 2020VISION / Common frog (Rana temporaria) leaping into a pond, controlled conditions, UK.

Michel Roggo / Grayling (Thymallus thymallus) on spawning ground. Lac de Fenetre, alpine lake, 2456 m above sea level, Canton of Valais, Switzerland, June.

Michel Roggo / Sockeye / Red Salmon (Oncorhynchus nerka) on spawning migration. Adams River, British Columbia, Canada, October.

Juan Carlos Munoz / Bearded vulture (Gypaetus barbatus) chasing Thick-billed Raven (Corvus crassirostris) in flight. Simien National Park, Ethiopia, Africa.

XI ZHINONG / Grey Wolves (Canis lupus) hunting a wounded Tibetan wild ass (Equus kiang) Kekexili, Qinghai, China, December.

Kristel Richard / Portrait of a grey Andalusian stallion, in Ciutadella, Menorca, Spain.

Steven Kazlowski / Dall sheep (Ovis dalli) solitary animal at sunset with a storm moving across the sky, Denali National Park, interior Alaska, USA

Claudio Contreras / Heermann's Gull (Larus heermanni) in flight, Ensenada, Baja California Peninsula, Mexico, December.

Paul D Stewart / Illustration of trilobuite (Paradoxides bohemicus), Plate 1. From 'Organization of the Trilobites' by Hermann Burmeister, appearing in the Ray Society translation of his work published in 1846. The Il...

Jordi Chias / Green Turtle, (Chelonia mydas), Swimming over volcanic sandy bottom, Armenime cove, South Tenerife coast, Canary Islands, Spain, Atlantic Ocean, May 2011, Commended in Wildlife Photographer of the Yea...

Eric Medard / Bechstein's Bat (Myotis bechsteinii) in flight at night, mouth open to emit echolocating calls. France, Europe, September.

Juan Carlos Munoz / Aerial view of flock of Flamingos flying over the salt lake, Lake Natron, Rift Valley, Tanzania, August 2009

Juan Carlos Munoz / Aerial view of flock of Flamingos flying over the salt lake, Lake Natron, Rift Valley, Tanzania, August 2009

Carol Walker / Cowboy galloping through snow, wearing thick sheepskin coat, Wyoming, USA, February 2012, model released

Carol Walker / White horses of the Camargue, herd running through water, Camargue, Southern France

Richard Du Toit / Springbok (Antidorcas marsupialis) silhouetted on the horizon. Namaqualand, South Africa, December.

Erlend Haarberg / Norway lemming (Lemmus lemmus) swimming, Sweden, May

Ernie Janes / Greylag geese (Anser anser) in flight with paraglider behind, Cley bank, Norfolk, UK

Andy Sands / Long Water Measurer (Hydrometra stagnorum) drinking by piercing surface tension with mouthparts. Captive, UK, July.

Marie Read / Western grebe (Aechmophorus occidentalis) pair swimming, with reflections, Bolsa Chica Ecological Reserve, California, USA, February

Eric Baccega / Muskrat (Ondatra zibethicus) walking on snow, Quebec, Canada

Roy Mangersnes / Black-legged kittiwakes (Rissa tridactyla) in flight during evening snowfall, Northern Nordaustlandet, Svalbard, Norway, August

Roy Mangersnes / Herring gull (Larus argentatus) in flight with others on water, Norway, July

Roy Mangersnes / Two Barnacle geese (Branta leucopsis) in flight, Spitsebergen, Svalbard, Norway, August

Mike Read / Cranes (Grus grus), imprinted from hatching to fly with a microlight. Hampshire County Show near Brockenhurst, Hampshire, England, July 2008.

Gerrit Vyn / Dall sheep (Ovis dalli) yearling lamb running across alpine tundra, Denali National Park, Alaska, USA

Nick Turner / Couple with child in seat cycling in the Cotswolds at Guiting Power, Gloucestershire, UK, August 2011. Model released

Paul Hobson / Birch Sawfly (Cimbex femoratus) feeding on a leaf. August, UK.

Paul Hobson / Feral Pigeon / Rock Dove (Columba livia) on city street. Sheffield, UK.

Alex Mustard / Yellowhead wrasse (Halichoeres garnoti) male performing a courtship dance to attract females, East End, Grand Cayman, Cayman Islands, British West Indies, Caribbean Sea.

Alex Mustard / Mirrorwing flying fish (Hirundichthys speculiger) mirrored and swimming beneath the surface at night, between Bahamas and Gulf Stream, West Atlantic Ocean, Florida, USA.

Carol Walker / Horse in snow storm with shed in background, USA

George Sanker / Grizzly bear (Ursus arctos horribilis) walking, Yellowstone National Park, Wyoming, USA, October

Edwin Giesbers / European mink (Mustela lutreola) swimming, captive, Germany, Critically endangered

Laurie Campbell / Black-headed gulls (Larus ridibundus) robbing Puffin (Fratercula arctica) of catch, Farne Islands, Northumberland, England, June

Petra Wegner / Cavalier King Charles Spaniel, ruby, running out of the sea wearing a lifejacket / life vest.

Sergey Gorshkov / Willow grouse / ptarmigan (Lagopus lagopus) pair in flight, Agapa River, Taimyr Peninsula, Siberia, Russia

Inaki Relanzon / Whitsunday Islands, aerial view with hydroplane landing, Great Barrier Coral Reef, Queensland, Australia, October 2011.

Inaki Relanzon / Trashumancia / seasonal migration, large herd of moving livestock, French Pryrenees, Spain, June 2011.

Brent Stephenson / Gentoo Penguins (Pygoscelis papua) porpoising on the way back to the breeding colony. Port Lockroy, Antarctic Peninsula, Antarctica, January.

Brent Stephenson / Flock of Cape petrels (Daption capense capense) in flight against the sea. Against the sun. South Shetland Islands, Antarctica, January.

Brent Stephenson / Flock of Cape Petrels (Daption capense capense) in flight, heading away against the sky. South Shetland Islands, Antarctica, December.

Brent Stephenson / New Zealand Albatross (Diomedea antipodensis) in flight, showing upperwing, wingtip touching the sea surface. Off Kaikoura, Canterbury, New Zealand, October.

Brent Stephenson / Dusky Dolphins (Lagenorhynchus obscurus) porpoising as they swim beside a boat. Off Kaikoura, Canterbury, New Zealand, October.

Brent Stephenson / Male Australasian Gannet (Morus serrator) flying with seaweed for use as nesting material. Cape Kidnappers, Hawkes Bay, New Zealand, September.

Todd Mintz / Sockeye salmon (Oncorhynchus nerka) split level view of annual spawning run, Adams River, British Columbia, Canada, October

Todd Mintz / Sockeye salmon (Oncorhynchus nerka) split level view of annual spawning run, Adams River, British Columbia, Canada, October

Todd Mintz / Great Horned owl, (Bubo virginianus) in flight, Regina, Saskatchewan, Canada, March

Todd Mintz / Narwhal (Monodon monoceros) diving sequence 1/4, Pond Inlet, Nunavut, Baffin Island, Canadian Arctic, June

Robin Chittenden / Mute swan (Cygnus olor) on water, Wheatfen, Norfolk, UK, November

Luiz Claudio Marigo / Aterro do Flamengo, a park designed by Roberto Burle Marx, with Rio de Janeiro city and the Sugar Loaf mountain in the background, Rio de Janeiro State, Brazil, May 2011

Philippe Clement / The harbour at Le Croisic, Loire-Atlantique, Pays-de-la-Loire, France September 2011

Markus Varesvuo / Spotted Redshank (Tringa erythropus) in flight, Utsjoki Finland June

Markus Varesvuo / Smew female (Mergus albellus) Kuusamo Finland May

Markus Varesvuo / Magpie (Pica pica) in flight, Sweden October

Markus Varesvuo / Dunlin (Calidris alpina) flock in flight, Finland July

Pete Oxford / Caribbean reef shark (Carcharhinus perezi) Jardines de la Reina National Park, Cuba

Pete Oxford / Vintage car on road in Havana, capital of Cuba, Caribbean 2011

Pete Oxford / Caribbean trumpetfish (Aulostomus maculatus) Bonaire, Netherlands Antilles, Caribbean

Luis Quinta / Baskian viper (Vipera seoanei) Geres National Park, Portugal, September.

Pedro Narra / Common dolphins ( Delphinus delphis) swimming below the surface, Atlantic Ocean, Portugal, October

Edwin Giesbers / Two Scarlet macaws (Ara macao) flying, captive

Chadden Hunter / King penguins (Aptenodytes patagonicus) walk along beach, South Georgia. Taken on location for BBC Frozen Planet series, 2008

Chadden Hunter / Emperor penguins (Aptenodytes forsteri) explode out of the water, returning to breed at Cape Washington, Antarctica. Taken on location for BBC Frozen Planet series, 2009

Chadden Hunter / Emperor penguins (Aptenodytes forsteri) males dive through openings in the sea ice to take their first swim in over 4 months as they have spent the bleak winter incubating their eggs. Cape Washington,...

Chadden Hunter / King penguins (Aptenodytes patagonicus) crossing water to reach breeding site, South Georgia. Taken on location for BBC Frozen Planet series, 2008

Thomas Rabeil / Flock of Marabou Stork (Leptoptilos crumeniferus) taking flight at seasonal waterhole, seen from above. Sahelo-Sudanese Biome, W National Park (UNESCO, IUCN & RAMSAR), Niger. Aerial census, May 2011.

John Abbott / Eastern Green june beetle (Cotinis nitida) in flight, Texas, USA, October

John Abbott / Snakefly (Agulla sp) in flight, Wild Basin Preserve, Texas, USA, April

Loic Poidevin / Redshank (Tringa totanus) walking on mud. Breton Marsh, French Atlantic Coast, November.

Juan Carlos Munoz / Jersey tiger moth (Euplagia quadripunctaria) large numbers flying to and resting on Sweetgum trees (Liquidambar orientalis) in the Petaloudes Valley, Rhodes island, Greece, August 2011

Gary K. Smith / Domestic sheep (Ovis aries) being moved from pen at livestock market after auction, Hawes Mart, Yorkshire, UK, September 2011

Nick Upton / Great capricorn / Great oak longhorn beetle (Cerambyx cerdo), one of the largest beetles in Europe, on tree trunk, Lesbos (Lesvos), Greece, June.

Kristel Richard / A purebred Lusitano stallion galloping in the arena at the Companhia das Lezírias Stud Farm, Samora Correia, Santarém, Portugal, July 2011

Kristel Richard / A purebred Lusitano colt (with his mane and tail cut to mark his first year) trotting in the arena at the Companhia das Lezírias Stud Farm, Samora Correia, Santarém, Portugal, July 2011

Martin Gabriel / Four Griffon Vultures (Gyps fulvus) in flight, silhouetted overhead. Monfrague National Park, Extremadura, Spain, March.

Doug Perrine / Hawaiian / Gray's / Long-Snouted Spinner Dolphins (Stenella longirostris longirostris) with mother and young calf. Keauhou, Kona, Hawaii, Central Pacific Ocean.

John Abbott / Ichneumon wasp (Compsocryptus sp.?) in flight, Williamson county, Texas, USA, November

Juan Carlos Munoz / White Tailed Sea Eagles (Aliaeetus albicilla) flying above snow with a Raven (Corvus corax) landed. Scandinavia, Europe, April.

Ben Osborne / Aerial view of African elephants (Loxodonta africana)moving through wetland in the Okavango delta, Northern Botswana, taken on location for BBC Planet Earth series, October 2005

Nick Garbutt / Silhouette of Red-fronted brown lemur (Eulemur rufus) leaping across gap in trees by moonlight, cathemeral behaviour, Ranomafana National Park, south east Madagascar, November

Michael W. Richards / Domesticated Indian elephants (Elephas maximus) being walked to forest at sunset, Pench Tiger Reserve, Madhya Pradesh, India

Michael W. Richards / Domesticated Indian elephants (Elephas maximus) being walked to forest at end of the working day, Pench Tiger Reserve, Madhya Pradesh, India

Sven Zacek / Roe Deer (Capreolus capreolus) herd travelling through snow and fog. Virumaa, Estonia, Europe, March.

Sven Zacek / House Martin (Delichon urbicum) in flight with invertebrates in its beak. Estonia, Europe, July.

Sven Zacek / Barn Swallow (Hirundo rustica) in flight. Estonia, Europe, June.

Sven Zacek / Geese on their spring migration above dawn lit clouds. Estonia, Europe, May.

Sven Zacek / Ural Owl (Strix uralensis) in flight over snow. Virumaa, Estonia, Europe, February.

Sven Zacek / Honey Bees (Apis mellifera) flying around the entrance to their (domestic) hive. Estonia, Europe, June.