Benjamin Barthelemy,Benjamin Barthelemy / Ibex (Capra Ibex) fighting, Aiguilles Rouges National Nature Reserve, Haute-Savoie, France, June

Daniel Heuclin / Horned viper (Vipera ammodytes) in natural setting, Turkey.

Stefan Christmann / Landscape of Emperor penguin (Aptenodytes forsteri) habitat with sun halo, Atka Bay, Queen Maud Land, Antarctica. October

Eric Baccega / Giant panda (Ailuropoda melanoleuca) female, Huan Huan, out in her enclosure in mist, Captive at Beauval Zoo, Saint Aignan sur Cher, France. The mist is created artificially by machine, in order to cr...

Eric Baccega / Giant panda (Ailuropoda melanoleuca) female, Huan Huan, out in her enclosure in mist, Captive at Beauval Zoo, Saint Aignan sur Cher, France The mist is created artificially by machine, in order to cr...

Andy Rouse / Great crested grebe (Podiceps cristatus) swimming, backlit with morning light forming a bokeh affect on the water, Wales, UK, February. Highly commended in the Habitat Category of the BWPA (British W...

Floris van Breugel / Beavertail cactus (Opuntia basilaris) in otherwise stark desert, Alabama Hills, Owen's Valley, California, USA May

Floris van Breugel / First rays of golden sunshine hit Giant Sequoias (Sequoiadendron giganteum) covered in a winter blanket of snow and frost, Grant Grove, Sequoia / Kings Canyon National Park, California, USA November

Wim van den Heever / Tranquil landscape with African elephant (Loxodonta africana) and rays of sunlight at sunrise, Masai Mara, Kenya.

Andres M. Dominguez / False death cap mushroom (Amanita citrina) Los Alcornocales Natural Park, Cortes de la Frontera, Malaga, Spain

Andres M. Dominguez / European stonechat (Saxicola rubicola) male perched with mountainous habitat behind, Sierra de Grazalema Natural Park, southern Spain, February.

Andres M. Dominguez / Fire salamander (Salamandra salamandra) in a Beech forest (Fagus sylvatica) Saja Besaya Natural Park, Saja, Cantabria, Spain, August.

Andres M. Dominguez / Portuguese oak tree (Quercus faginea) covered in moss, Los Alcornocales Natural Park, southern Spain, November.

Sebastian Kennerknecht / Agricultural land surrounded by natural habitat, Santa Cruz Mountains, Monterey Bay, California

Wim van den Heever / Quiver tree forest (Aloe dichotoma) at night with stars and the milky way, Keetmanshoop, Namibia.

Wim van den Heever / Quiver tree forest (Aloe dichotoma) at night with stars and the Milky Way, Keetmanshoop, Namibia.

Radomir Jakubowski / Red fox (Vulpes vulpes) standing in snowfall, Grand Teton National Park, Wyoming, USA, February. Winner of the NPL Best Single Image award in the Terre Sauvage Nature Images Awards competition 2015.

Juergen Freund / Split level view of coral reef, Komodo National Park, Flores, Lesser Sunda Islands, Indonesia. Second Place in the Portfolio Award of the Terre Sauvage Nature Images Awards Competition 2015.

Alex Mustard / Colourful reef scene with Coral grouper (Cephalopholis miniata), map puffer (Arothron mappa) and Panda butterflyfish (Chaetodon adiergastos). Tank Reef, Fiabacet Islands, Misool, Raja Ampat, Indonesia...

Mark MacEwen / Northern royal albatross (Diomedea sanfordi), early pairing behaviour, Taiaroa Head, Otago Peninsula, New Zealand. Endangered.

Mark MacEwen / Snare's island crested penguin (Eudyptes robustus) colony on the coast, high angle view, Snares Island, New Zealand.

Mark Hamblin / Mountain hare (Lepus timidus) in white winter coat stretching - in snowy habitat, Scotland. Highly Commended in the Mammals category of the GDT European Photographer of the Year Competition 2015.

Cyril Ruoso / Queen of the Andes (Puya raymondii) Cordillera Blanca Massif, Andes, Peru, November.

Cyril Ruoso / South american sea lion (Otaria byronia) and American fur seal (Arctocephalus australis) hauled out on the shore of Punta Coles Reserve, Peru.

Alex Hyde / Red Deer herd (Cervus elaphus) moving over mountain ridge in heavy snow. Cairngorms National Park, Scotland. January. Highly Commended in the Habitat category of the British Wildlife Photography Award...

Staffan Widstrand / Wild boar (Sus scrofa) in old mixed conifer and broadleaf forest, Punia Forest Reserve, Lithuania, May.

Sven Zacek / Autumn trees on the Rapa River Delta, aerial view, Rapadalen Sarek National Park, Sweden, September 2009.

Konrad Wothe / View up into canopy with sun rays shining through the leaves, lowland rainforest, Panguana Reserve, Huanuco province, Amazon basin, Peru.

Pete Oxford / Guanaco (Lama guanaco) with Cordiera del Paine in back. Torres del Paine National Park, Patagonia, Chile.

Cyril Ruoso / European hedgehog (Erinaceus europaeus) at dusk, Picardy, France. Non-ex

David Pattyn / Alpine ibex (Capra ibex) two adult males in mountain landscape. Alps, Aosta Valley, Gran Paradiso National Park, Italy. September.

Steve O. Taylor (GHF) / Loss of topsoil in deforested land, Central Kalimantan, Indonesian Borneo. June 2010.

Krista Schlyer / Yellow pitcher (Sarracenia flava) plants in bog, growing in fire-maintained landscape, Conecuh National Forest, Alabama, USA. May 2014.

Stephen Dalton / View to South Downs over farmland, Sussex, England, UK. October.

Kristel Richard / Two wild Przewalski / Takhi Horse (Equus ferus przewalskii) stallions play fighting Hustai National Park, Tuv Province, Mongolia. Endangered species. September.

Jussi Murtosaari / Apollo butterfly (Parnassius apollo) in flight, Foglo, Ahvenanmaa / Aland Islands, Finland, July.

Tui De Roy / Erect-crested penguin (Eudyptes sclateri) pair. Antipodes Island, New Zealand Sub-Antarctic Islands. Endemic to Antipodes and Bounty Islands. Endangered species.

David Tipling / Red campion (Silene dioica) flowering in woodland amongst Bluebells and Greater stitchwort. Thursford Wood, North Norfolk, UK, May.

Floris van Breugel / Two Guanacos (Lama guanicoe) at edge of lake with Cuernos del Paine beyond. Torres del Paine National Park, Chile, June 2014.

David Pattyn / Heathland landscape at sunset, Kampina Nature Reserve, Netherlands. August 2006.

Frederique Olivier / Adelie penguins (Pygoscelis adeliae) preening at edge of ice, Antarctica.

Enrique Lopez-Tapia / Graphite mine near the Andasibe-Mantadia National Park, one of the environmental problems affecting the ecosystem of the rainforest. Madagascar, March 2005.

Jack Dykinga / Male Andean Condor (Vultur gryphus) flying amongst the clouds, Los Cuernos Peaks, Torres del Paine, Chile. March.

Denis-Huot / Wildebeest (Connochaetes taurinus) herd below stormy sky during migration, Masai-Mara game reserve, Kenya.

Denis-Huot / Coalition of male Lions (Panthera leo) in grassland before storm, Masai-Mara game reserve, Kenya.

Pascal Kobeh / Group of California sea lions (Zalophus californianus) swimming in kelp forest (Macrocystis pyrifera), California, USA. Pacific ocean.

Ben Cranke / Fin whale (Balaenoptera physalus) surfacing with mountain landscape, Liefdefjorden, Svalbard, Norway. July 2011.

Steve O. Taylor (GHF) / Cut trees in the Waka forest, Gabon, Central Africa, 2008-2009.

Richard Robinson / Hector's dolphin (Cephalorhynchus hectori) Akaroa Harbour, South Island, New Zealand, November. Editorial use only.

Will Burrard-Lucas / Giant Panda (Ailuropoda melanoleuca) young male, Qinling Mountains, China, April.

Will Burrard-Lucas / Big-headed mole rat (Tachyoryctes macrocephalus) appearing from its hole, Bale Mountains National Park, Ethiopia. Taken with remote camera.

Michel Roggo / Underwater river scene with freshwater plants and Tetra fish, Aquario Natural, Rio Baia Bonito, Bonito area, Serra da Bodoquena (Bodoquena Mountain Range), Mato Grosso del Sul, Brazil November/Decembe...

Radomir Jakubowski / Alpine ibex (Capra ibex) male on mountainside at sunrise, Bernese Alps, Switzerland, July. Received honourable mention in the Big Picture Competition awards 2014. Nominated in the Melvita Nature Image...

Dong Lei / Blood pheasants (Ithaginis cruentus) foraging in habitat, one looking at the camera, Meli Snow Mountain National Park, Yunnan Province, China, January. Endemic. Taken with remote camera.

Dong Lei / Himalayan goral (Naemorhedus goral) in forest, Tangjianhe National Nature Reserve, Sichuan Province, China. April.

Anup Shah / Lion (Panthera leo) cub aged about 2 years approaching with curiosity - remote wide angle perspective . Masai Mara National Reserve, Kenya. Taken with remote wide angle camera.

Anup Shah / Cape buffalo (Syncerus caffer) young male approaching with curiosity. Masai Mara Nationa Reserve, Kenya. Taken with remote wide angle camera.

Michel Roggo / Anavilhanas Archipelago in the Rio Negro, and flooded forest or 'Varzea forest' Amazonas, Brazil. February 2011

Ann & Steve Toon / Namaqua chameleon (Chamaeleo namaquensis), Namib Desert, Namibia, April

Ben Hall / Stunted oak woodland covered in moss, Wistman's Wood, Devon, UK. August. Highly Commended in the Wild Woods category of the British Wildlife Photography Awards (BWPA) competition 2013.

Roland Seitre / Natural pool and Cook's pine trees (Araucaria columnaris), Oro, Ile des Pins / Isle of Pines, South Province, New Caledonia.

Daniel Heuclin / Malayan Tiger (Panthera tigris jacksoni) in river, Malaysia. Captive in natural setting.

Franco Banfi / Piraputanga (Brycon hilarii) reflecting on the water surface, Aquirio Natural, Bonito, Mato Grosso do Sul, Brazil

Theo Bosboom,Theo Bosboom / Gulls, predominantly Iceland gulls (Larus glaucoides) flying above stormy ocean near Vik, Iceland, March 2012.

Sandesh Kadur / Clouded leopard (Neofelis nebulosa) walking along fallen tree trunk, Assam, India, captive. Vulnerable species.

Juan Carlos Munoz / Walia ibex (Capra walie) herd, Simien Mountains National Park, Ethiopia

Vladimir Medvedev,Vladimir Medvedev / Amur / Siberian Tiger (Panthera tigris altaica) female in the wild, on a hillside, Lazovskiy zapovednik / Lazo Reserve protected area, Primorskiy krai, Far Eastern Russia, February 2012

Paul Harcourt Davies,Paul Harcourt Davies / Elder flower orchid (Dactylorhiza sambucina) in its two colour forms, Campo Imperatore, Gran Sasso, Appennines, Abruzzo, Italy, May

Paul Harcourt Davies,Paul Harcourt Davies / Yellow tulip (Tulipa australis) in flower, above Piano Grande, Sibillini, Appennines, Umbria, Italy, May 2011

Paul Harcourt Davies / Spring Crocus (Crocus vernus) in flower in snow, Campo Imperatore, Gran Sasso, Appennines, Abruzzo, Italy, May

Benjamin Barthelemy,Benjamin Barthelemy / Alpine ibex (Capra ibex ibex) in front of the Mont Blanc massif, seen from the Aiguilles Rouges (Red Peaks) Regional Natural Park, Haute-Savoie, France, June.

Erlend Haarberg,Erlend Haarberg / Aerial view down river to the east of the Hofsjakull glacier and west of the Sprengisandur upland desert. Innumerable rivulets flow from beneath the glacier in this area of tundra which abounds in pon...

David Woodfall / Sessile oak (Quercus petraea) tree with sunlight behind, ancient semi natural woodland in Gilfach Nature Reserve, Radnorshire Wildlife Trust, Powys, Wales, UK May

Sven Zacek / Lowland bog landscape seen from the air. South Estonia, Europe, October 2010.

Orsolya Haarberg / Red Knots (Calidris canutus) during migration rest on coastal rocks as waves wash across the coastline. The Snæfellsnes peninsula, Iceland, May. Highly commended in the Birds category of the GDT 2011...

Luiz Claudio Marigo / A field of Yellow 'Frailejon' (Espeletia grandiflora). Paramo habitat (Andean high altitude fields) Chingaza Natural National Park, municipality of La Calera, Cundinamarca Department, Colombia.

Chris Gomersall / 2020VISION / RSPB reserves staff thinning woodland encroaching onto heath at Westleton Heath (Natural England), Suffolk, UK, February 2011.

Juergen Freund / Common death adder (Acanthophis antarcticus) well camouflaged in its natural habitat. Queensland, Australia, February

Alex Mustard / Southern stingray (Hypanus americanus) swimming over ripples on sandbar, Grand Cayman, Cayman Islands.

Jouan & Rius / Smoke from bush fire in Pilbara region, Western Australia. August 2009

Jouan & Rius / Wave rock, an eroded granite plateau, Hyden area, Western Australia. August 2009

Jouan & Rius / Wave rock, an eroded granite plateau, Hyden area, Western Australia. August 2009

Tim Laman / Praying mantis (Mantodea) green leaf mimic type walking along a mossy branch in lowland rainforest, Gunung Palung National Park, Borneo, West Kalimantan, Indonesia

Alan Watson / Frond pattern of tree ferns (Cyathea arborea) in lowland tropical rainforest at 420 metres, Loma Quita Espuela Scientific Reserve, Dominican Republic.

Wild Wonders of Europe / Oxford / Wild Iberian lynx (Lynx pardinus) female emerging from undergrowth, Sierra de Andjar Natural Park, Mediterranean woodland of Sierra Morena, north east Jan Province, Andalusia, Spain, April 2009, Criti...

Laurie Campbell / Bluebells (Hyacinthoides non-scripta) flowering in oakwood, Perthshire, Scotland, UK, May

Angelo Gandolfi / Sunset at Salto del Roldan canyon, a nesting place for birds of prey. Sierra de Guara, Aragon, Spain, 2007.

Mark Bowler / Tropical rainforest habitat near the Yavari river, Peru

Mark Bowler / Tropical rainforest habitat near the Yavari river, Peru

Andy Rouse / Blue Tit (Parus caeruleus) perched in snow, Wales, UK, January

Wild Wonders of Europe / Oxford / Wild Iberian lynx (Lynx pardinus) female, one year walking, Sierra de Andjar Natural Park, Mediterranean woodland of Sierra Morena, north east Jan Province, Andalusia, Spain, April 2009, Critically en...

Luiz Claudio Marigo / Maned wolf (Chrysocyon brachyurus) in its natural habitat, at sunset, the grasslands of the Brazilian Cerrado, Serra da Canastra National Park, Minas Gerais State, Brazil.

Jack Dykinga / Gopher snake (Pituophis sp) amongst Lechuguilla (Agave lechuguilla) and prickly pear cacti (Opuntia macrocentra), Maderas del Carmen Natural Reserve, Mexico

Piotr Naskrecki / Splendid Leaf Frog (Agalychnis calcarifer) a rarely seen species associated with natural forest gaps, Costa Rica

Tui De Roy / King Vulture (Sarcoramphus papa) in natural arid forest habitat, Cerro Chaparri, Lambayeque Province, Peru