Bence Mate / European rabbits (Oryctolagus cuniculus) fighting each other, Kiskunsag National Park, Hungary. June. Winner, 2019 Cadiz Photo Nature Competition.

Ann & Steve Toon / Meerkats (Suricata suricatta) on the move, Kgalagadi Transfrontier Park, Northern Cape, South Africa, January. Second place in the Mammals Category of the GDT European Nature Photographer of the Year...

Gary Bell/ Oceanwide / Sea Goldie (Pseudanthias squamipinnis) school in coral reef with feather stars, Great Barrier Reef, Queensland, Australia

Oscar Dewhurst / Fallow deer (Dama dama) walking across frost-covered playing fields at dawn, with tower blocks of London in the background. Richmond Park, London, UK. December

Oscar Dewhurst / Red deer (Cervus elaphus) stag roaring, silhouetted under trees at sunrise. Bushy Park, London, UK. September

Marie Lochman / Biodiversity of flora in heath / Kwongan habitat, Wheatbelt, Frank Hann National Park, November 2013

Yashpal Rathore / Bengal tiger (Panthera tigris tigris) walking at night, with monsoon clouds and lightning. Kanha National Park, Central India. Dominant male (T29)

SCOTLAND: The Big Picture / Ancient Scots pine (Pinus sylvestris) amongst flowering Common heather / Ling (Calluna vulgaris). Rothiemurchus Forest, Cairngorms National Park, Scotland, UK.

Christophe Courteau / Mountain gorilla (Gorilla beringei beringei) blackback, juvenile male demonstrating power. Volcanoes National Park, Rwanda.

Christophe Courteau / Mountain gorilla (Gorilla beringei beringei) blackback, juvenile male demonstrating power. Volcanoes National Park, Rwanda.

Ingo Arndt / Guanaco (Lama guanicoe) herd, Torres del Paine National Park, Patagonia, Chile

Ingo Arndt / Mountain Lion (Puma concolor) hunting Guanaco (Lama guanicoe) male, Torres del Paine National Park, Patagonia, Chile, sequence 4 of 12. Highly commended in the Wildlife category of the Nature?s Best P...

Tui De Roy / Blue-footed booby (Sula nebouxii) landing. Punta Vicente Roca, Isabela Island, Galapagos.

Bence Mate / Great white egret (Ardea alba) group of three in winter, Kiskunsag National Park, Hungary. Winner of the Bird Category of Nature Photographer of the Year Awards 2018.

Jeff Foott / Wild California condor (Gymnogyps californianus) near San Pedro Martir National Park, Northern Baja California, Mexico.

Chien Lee / Limestone pinnacles, Tsingy de Bemaraha National Park, Madagascar

Jeff Foott / Maple (Acer sp) leaves and needles in autumn, Acadia National Park, Maine

Jack Dykinga / Western sword fern (Polystichum munitum) and Big leaf maple (Acer macrophyllum) trees that are covered with moss, Hoh Temperate Rainforest, Olympic National Park, Washington, USA. June 2017.

Claudio Contreras / Aerial view of Espiritu Santo Island National Park, Sea of Cortez (Gulf of California), Mexico, June, Honorary Mention in the third national contest 'Visions of our Nature' organised by Mexico...

Claudio Contreras / Prickly pear (Opuntia sp.) growing on coastline, Socorro Island, Revillagigedo Archipelago National Park (Socorro Islands), Pacific Ocean, Western Mexico, November, Honorary Mention in the third natio...

Paul Bertner / Blue-legged Chameleon (Calumma crypticum) curled tail, Ranomafana National Park, Madagascar

Tony Phelps / Cape cobra (Naja nivea) spreading its hood, retreats into hole, De Hoop Nature Reserve, Western Cape, South Africa, January.

Tony Phelps / Cape cobra (Naja nivea) emerging from a Cape gerbil (Gerbilliscus afra) burrow, hunting and smelling with its tongue, De Hoop Nature Reserve, Western Cape, South Africa, January.

Tony Phelps / Cape cobra (Naja nivea) hunting, chasing a Cape gerbil (Gerbilliscus afra), De Hoop Nature Reserve, Western Cape, South Africa, December.

Tony Phelps / Puff adder (Bitis arietans) moving into ambush position under a bush, DeHoop Nature Reserve, Western Cape, South Africa, January.

Tony Phelps / Chacma baboon (Papio ursinus) troop exiting den, DeHoop Nature Reserve, Western Cape, South Africa, June.

Ingo Arndt / Floodplain with Pandanus palms under storm clouds, Kakadu National Park, Northern Territory, Australia, December 2010

Ingo Arndt / View of Badlands National Park, with shadows cast between mountain peaks, and sediment layers visible, South Dakota, USA. September 2009.

Ingo Arndt / View of Badlands National Park, with shadows cast between mountain peaks, and sediment layers visible, South Dakota, USA. September 2009.

Ingo Arndt / Grizzly Bear (Ursus arctos horribilis) digging for clams on tidal flats, Lake Clark National Park, Alaska.

Ingo Arndt / Gelada baboon troop on rock {Theropithecus gelada} Simien Mt NP, Ethiopia

Richard Du Toit / Black Wildebeest (Connochaetes gnou), Rietvlei Nature Reserve, South Africa

SCOTLAND: The Big Picture / Osprey (Pandion haliaetus) male flying with talons outstretched, Glenfeshie, Cairngorms National Park, Scotland, UK, May.

Sandra Bartocha / European beech tree (Fagus sylvatica). in woodland, Serrahn, Muritz-National Park, World Natural Heritage site, Germany, Europe. October 2015.

Sandra Bartocha / Sun rays shining through European beech woodland (Fagus sylvatica). Serrahn, Muritz-National Park, World Natural Heritage site, Germany, Europe. November 2015.

Chris & Monique Fallows / Impala (Aepyceros melampus) herd at dawn, Mana Pools National Park, Zimbabwe, October.

Vincent Grafhorst / Lava in volcanic crater, Mount Nyiragongo, Virunga National Park, Democratic Republic of the Congo

Richard Du Toit / Burchell's Zebra (Equus burchellii) pair nuzzling, Rietvlei Nature Reserve, South Africa

Richard Du Toit / Burchell's Zebra (Equus burchellii) females nuzzling, Rietvlei Nature Reserve, South Africa

Richard Du Toit / Cape Buffalo (Syncerus caffer) herd chasing away Cheetah (Acinonyx jubatus), Rietvlei Nature Reserve, South Africa

Richard Du Toit / Burchell's Zebra (Equus burchellii) mother and foal, Rietvlei Nature Reserve, South Africa

Richard Du Toit / Burchell's Zebra (Equus burchellii) foals, Rietvlei Nature Reserve, South Africa

Richard Du Toit / Burchell's Zebra (Equus burchellii) parents tending to foal, Rietvlei Nature Reserve, South Africa

Andres M. Dominguez / Mirror orchid (Ophrys speculum) in the Sierra de Grazalema Natural Park, El Bosque, southern Spain, April.

Andres M. Dominguez / Frozen Elmleaf blackberry (Rubus ulmifolius) in a small pond, Sierra de Grazalema Natural Park, southern Spain, January.

Orsolya Haarberg / Snow covered Norway spruce (Picea abies) trees at the treeline, Stubba Nature Reserve, Laponia, Sweden, February 2013.

Wim van den Heever / Blue wildebeest (Connochaetes taurinus) herd silhouetted against the rising sun with clouds in the background. Greater Kruger National Park, South Africa

Wim van den Heever / Snow Leopard (Panthera uncia) climbing up mountain slope, Hemis National Park, India, February.

Wim van den Heever / Blue sheep (Pseudois nayaur) Hemis National Park, Jammu and Kashmir, India.

Erlend Haarberg / Photographer silhouetted on Mt Luottotjakka overlooking the Arjep Luottojekna and Nuortap glaciers at sunset, Sarek National Park, World Heritage Laponia, Swedish Lapland, Sweden, August 2013.

Jeff Vanuga / Old Faithful Geyser erupting at -30 Celsius in winter. Upper Geyser Basin, Yellowstone National Park, Wyoming, USA, January 2015.

Cheryl-Samantha Owen / Common or plains zebra (Equus quagga burchelli) grazing in bushTarangire National Park, Northern Tanzania.

Wim van den Heever / Leopard (Panthera pardus) hunting Springbok (Antidorcas marsupialis) Etosha, Namibia, Sequence 4 of 5

Radomir Jakubowski / Red fox (Vulpes vulpes) standing in snowfall, Grand Teton National Park, Wyoming, USA, February. Winner of the NPL Best Single Image award in the Terre Sauvage Nature Images Awards competition 2015.

Juergen Freund / Split level view of coral reef, Komodo National Park, Flores, Lesser Sunda Islands, Indonesia. Second Place in the Portfolio Award of the Terre Sauvage Nature Images Awards Competition 2015.

Staffan Widstrand / Hippopotamus (Hippopotamus amphibius) bull emerging from water to charge, iSimangaliso Wetland Park, KwaZulu Natal National Park. South Africa. Specially commended in the Animal Portraits category of...

Bence Mate / White-tailed Eagle (Haliaeetus albicilla) flying, with Great egret (Ardea alba) in the foreground, Lake Csaj, Pusztaszer, Hungary. February. Winner of the Portfolio category of the Terre Sauvage Natur...

Thomas Marent / Black-headed Night Monkey (Aotus nigriceps) at night, Tambopata-Candamo Nature Reserve, Peru

Thomas Marent / Saddle-back Tamarin (Saguinus fuscicollis) mother with young, Tambopata-Candamo Nature Reserve, Peru

Thomas Marent / Black-faced Spider Monkey (Ateles chamek) juvenile calling, Tambopata-Candamo Nature Reserve, Peru

Alex Mustard / Harlequin sweetlips (Plectorhinchus chaetodonoides) cleaned by a pair of bluestreak cleaner wrasse (Labroides dimidiatus). South Atoll, Tubbataha Atolls, Tubbataha Reefs Natural Park, Palawan, Philipp...

Alex Mustard / Napoleon wrasse (Cheilinus undulatus) super male, patrolling coral, Ras Mohammed Marine Park, Sinai, Egypt. Red Sea

Alex Mustard / Group of predators hunting together, in an example of 'Nuclear foraging'. The Moray eel (Gymnothorax javanicus) flushes out prey, and is followed by a Napolean wrasse (Cheilinus undulatus) and a Bluef...

Fiona Rogers / Eastern chimpanzee (Pan troglodytes schweinfurtheii) female 'Gremlin' aged 40 years holding female baby aged 2, her granddaughter. Gombe National Park, Tanzania. Gremlin took the new born baby from he...

Suzi Eszterhas / Brown Bear (Ursus arctos) mother and 3-4 month cubs foraging for clams, Katmai National Park, Alaska, USA. June.

Suzi Eszterhas / Brown Bear (Ursus arctos) mother and 3-4 month cubs foraging for clams. Katmai National Park, Alaska, USA, June.

Tony Heald / Hanuman langur (Semnopithecus entellus) baby next to mother, Bandhavgarh National Park, India.

Nick Garbutt / Madagascar sunset moth (Chrysiridia rhipheus) resting on tree. Masoala Peninsula, north east Madagascar. Controlled conditions.

Tony Heald / Bengal tiger (Panthera tigris tigris) walking, Bandhavgarh National Park, India.

Tony Heald / Indian rhinoceros (Rhinoceros unicornis) mother and calf, rear view. Kaziranga National Park, Assam, India. Vulnerable species

Sharon Heald / African buffalo (Syncerus caffer) low angle portrait, Chobe National Park, Botswana.

Alex Hyde / Red Deer herd (Cervus elaphus) moving over mountain ridge in heavy snow. Cairngorms National Park, Scotland. January. Highly Commended in the Habitat category of the British Wildlife Photography Award...

Doug Perrine / Barjacks (Caranx ruber), Yellowtail snapper (Ocyurus chrysurus), and Sergeant major damselfish (Abudefduf saxatilis) swimming around wreck of bow gun on C-58 shipwreck off Puerto Morelos, near Cancun,...

Sven Zacek / Pink algae in fresh snow on the coast of the Baltic Sea, Lahemaa National Park, Northern Estonia. December 2012.

George Sanker / Great egret (Ardea alba) taking off with fish prey in mouth, Myakka River State Park, Florida, USA, March.

George Sanker / White-tailed deer (Odocoileus virginianus) female grazing in snow, Acadia National Park, Maine, USA. February.

George Sanker / North American river otter (Lontra canadensis) two in water with autumnal trees reflected in the water, Acadia National Park, Maine, USA, October.

Dong Lei / Himalayan honeybee (Apis laboriosa) Mount Namjagbarwa, Yarlung Zangbo Grand Canyon National Park, Nyingchi Prefecture, Tibet, China. October.

George Sanker / North American beaver (Castor canadensis). swimming in late evening light, Acadia National Park, Maine, USA., September.

George Sanker / Purple gallinule (Porphyrula martinica) in mangrove swamp. Everglades National Park, Florida, USA. March.

Pete Oxford / Guanaco (Lama guanaco) with Cordiera del Paine in back. Torres del Paine National Park, Patagonia, Chile.

Gabriel Rojo / Wild puma (Puma concolor) walking across rocks, profile, Torres del Paine National Park, Chile.

Gabriel Rojo / Wild puma (Puma concolor) walking across rocks, Torres del Paine National Park, Chile.

Sean Crane / Celebes Black Macaque (Macaca nigra) scratching head, Tangkoko Nature Reserve, Sulawesi, Indonesia

Richard Du Toit / Ostrich (Struthio camelus) male in threat display, Rietvlei Nature Reserve, South Africa

Richard Du Toit / Burchell's Zebra (Equus burchellii) mother and foal in tall grass, Rietvlei Nature Reserve, South Africa

Alex Hyde / Adder (Vipera berus) basking amongst bracken fronds. Peak District National Park, Derbyshire, UK. April.

Alex Hyde / Crane fly / Daddy long legs (Tipula paludosa) female covered in morning dew. Peak DIstrict National Park, Derbyshire, UK. September. Winner of the Hidden Britain Category of the BWPA Awards 2015.

Enrique Lopez-Tapia / Giant lobelia (Lobelia rhynchopetalum) flower spikes in mist, Sanetti Plateau, Bale Mountains National Park. Ethiopia, November 2014

Eric Baccega / Spirit Island, Maligne Lake, Jasper National Park, Rocky Mountains, Alberta, Canada, September 2009

Scott Leslie / Longleaf Pine (Pinus palustris) forest, Okefenokee National Wildlife Refuge, Georgia

David Pattyn / Alpine ibex (Capra ibex) two adult males in mountain landscape. Alps, Aosta Valley, Gran Paradiso National Park, Italy. September.

Cyril Ruoso / Bromeliad (Bromeliaceae) pair flowering, Pacaya Samiria National Park, Peru

Kevin Schafer / Rainbow over cliffs, Kalalau Valley, Na Pali Coast State Park, Kauai, Hawaii

Erlend Haarberg / Three-toed woodpecker (Picoides tridactylus) on tree trunk, Sarek National Park, Laponia World Heritage Site, Lapland, Sweden, September.