Christophe Courteau / Blue footed booby displaying {Sula nebouxii} Galapagos.

Andy Rouse / Polar Bear (Ursus maritimus) portrait in sea-ice landscape. Wide angle / fish-eye shot. Svalbard, Norway, September.

Richard Du Toit / Cattle Egret (Bubulcus ibis) on Burchell's Zebra (Equus burchellii), Rietvlei Nature Reserve, Gauteng, South Africa

Piotr Naskrecki / Scorpion (Brotheas sp) in defensive posture seen under ultraviolet light, Suriname

David Tipling / Tourist photographing King Penguins (Aptenodytes patagonicus) Fortuna Bay, South Georgia, November 2006. No release available.

Gavin Hellier / Buddhist monks relaxing at Ta Phrohm Temple, Angkor Wat, Siem Reap, Cambodia 2010. Model released.

Gavin Hellier / Two buddhist monks relaxing in the Bayon Temple, Angkor Wat, Siem Reap, Cambodia 2010. Model released.

Andrew Parkinson / Male Capercaillie (Tetrao urogallus) displaying at photographer Andy Parkinson whilst on assignment for 2020VISION, Cairngorms National Park, Scotland, UK, March.

Neil Aldridge / Meerkat (Suricata suricatta) standing and keeping watch while another excavates a burrow in soft sand before disappearing underground for the night. Makgadikgadi Pans, Botswana, April.

Neil Aldridge / Young Meerkat (Suricata suricatta) standing on its hind legs near the edge of Makgadikgadi Pans National Park, Botswana, April.

Paul Hobson / Small garden bumble bee (Bombus hortorum) frozen on small twig, covered in ice crystals, Sheffield, UK February 2011

Fiona Rogers / Celebes / Black crested macaque (Macaca nigra) baby less than 1 month old in mothers arms, Tangkoko National Park, Sulawesi, Indonesia.

Anup Shah / Celebes / Black crested macaque (Macaca nigra) sub-adult male sitting on a beach, Tangkoko National Park, Sulawesi, Indonesia.

Anup Shah / Celebes / Black crested macaque (Macaca nigra) female sitting in a tree with her baby aged less than 1 month, Tangkoko National Park, Sulawesi, Indonesia.

Anup Shah / Celebes / Black crested macaque (Macaca nigra) silhouette of juvenile sitting in a tree at coast, Tangkoko National Park, Sulawesi, Indonesia.

Wild Wonders of Europe / Widstrand / Pool Frog (Rana lessonae) sitting on White lily pad, Danube delta rewilding area, Romania

Mark Taylor / Tabby kittens, Stanley and Fosset, 3 months old, sitting and snuggling together.

Mark Taylor / Tabby kittens, Stanley and Fosset, 12 weeks old, walking together.

Denis-Huot / Masai Giraffe (Giraffa camelopardalis tippelskirchi) male and female (back). Masai-Mara Game Reserve, Kenya.

Denis-Huot / African Elephant (Loxodonta africana) standing on hind legs to reach leaves high in tree. Masai-Mara Game Reserve, Kenya.

Denis-Huot / African Elephant (Loxodonta africana) baby elephant its mother's trunk Amboseli National Park, Kenya.

Mark Taylor / Beagle dog, Bruce, with ginger kitten, Tom.

Mark Taylor / Russian Blue female cat with green eyes.

David Tipling / Grey Squirrel (Sciurus carolinensis) climbing on fence, St James Park, London, UK, October

David Pattyn / Southern tamandua / Lesser anteataer (Tamandua tetradactyla) standing up in a defensive posture, Southern Pantanal, Brazil

Linda Pitkin / 2020VISION / White clawed crayfish (Austropotamobius pallipes) underwater on riverbed, showing defensive posture, River Leith, Cumbria, England, UK, September 2012. Did you know? White clawed crayfish can live fo...

Luke Massey / 2020VISION / Male Pheasant (Phasianus colchicus) displaying, Hertfordshire, England, UK

Anup Shah / Western chimpanzee (Pan troglodytes verus) juvenile female 'Joya' aged 6 years using rocks as tools to crack open palm oil nuts, Bossou Forest, Mont Nimba, Guinea. December 2010.

Staffan Widstrand / Polar Bear (Ursus maritimus) reflection in water, Svalbard, Norway

Anup Shah / Western chimpanzee (Pan troglodytes verus) males 'Jeje' aged 13 years, 'Tua' aged 53 years and 'Peley' aged 12 years grooming, Bossou Forest, Mont Nimba, Guinea. January 2011.

Anup Shah / Western chimpanzee (Pan troglodytes verus) young male 'Peley' aged 12 years displaying, Bossou Forest, Mont Nimba, Guinea. January 2011.

Anup Shah / Western chimpanzee (Pan troglodytes verus) juvenile female 'Joya' aged 6 years sitting in the trees, Bossou Forest, Mont Nimba, Guinea. December 2010.

Juergen Freund / Spectacled flying fox (Pteropus conspicillatus) casualty of paralytic tick (Ixodes holocyclus) bite being cared for on site of colony by Jenny Mclean, North Queensland, Australia, November 2012

Alex Hyde / Highland cow on heather moorland, Curbar Edge, Peak District National Park, Derbyshire. August.

Markus Varesvuo / Ruff (Philomachus pugnax) two males displaying in territorial dispute, Liminka Finland June

Markus Varesvuo / Capercaillie (Tetrao urogallus) close up of male calling in the snow, Vaala Finland April

Markus Varesvuo / Two Sparrowhawks (Accipiter nisus) immature male and female in flight, male on left, Falsterbo, Sweden. September.

XI ZHINONG / Tibetan antelope (Pantholops hodgsonii) male, Kekexili, Qinghai, Tibetan Plateau, China, November

Kristel Richard / Lady riding a black Menorquin stallion, performing the bot or walking courbette of the Doma Menorquina, during the festival Mare de Deu de Gracia, in Mahon, Menorca, Spain 2012. Tourists are trying t...

Fabrice Cahez / Wild Cat (Felis silvestris) in mist. Vosges, France, August.

Mark Carwardine / Humpback whale lobtailing {Megaptera novaengliae} Dominican Republic, Caribbean

Mark Carwardine / Humpback whale lobtailing {Megaptera novaengliae} Dominican Republic, Caribbean

Andrew Parkinson / Red Squirrel (Sciurus vulgaris) adult reaching up. Cairngorms National Park, Scotland, UK, March.

Bert Willaert / Sparrowhawk (Accipiter nisus) being caught by a falconer using a Red-backed shrike (Lanius collurio) as bait, Georgia, September 2011.

Steven Kazlowski / Grizzly bear (Ursus arctos horribilis) young rare blonde (white) bear plays with a young dark coloured bear along the coast of Katmai National Park, Alaskan peninsula, USA

Steven Kazlowski / Lynx (Lynx lynx) adult in the snowy foothills of the Takshanuk mountains, South-East Alaska, USA

Steven Kazlowski / Crew bleeds a full deckload of Pacific cod fish (Gadidae family) before putting them in salt water tanks, on the fishing trawler Western Dawn, Gulf of Alaska, USA. No release available.

Claudio Contreras / American Flamingo (Phoenicopterus ruber) juveniles following adults, Ria Lagartos Biosphere Reserve, Yucatan Peninsula, Mexico, August.

Jeff Rotman / Common Thresher Shark (Alopias vulpinus) caught on gill net being hauled up onto boat, Huatabampo, Mexico, Sea of Cortez, Pacific Ocean. Model released. Model released.

Paul D Stewart / Male peacock (Pavo cristatus) displaying his ocellated tail feathers.

Anup Shah / Western lowland gorilla (Gorilla gorilla gorilla) female 'Mopambi' sitting with her playful infant 'Sopo' aged 18 months, Bai Hokou, Dzanga Sangha Special Dense Forest Reserve, Central African Republi...

Anup Shah / Western lowland gorilla (Gorilla gorilla gorilla) rear view of dominant male Silverback 'Makumba' aged approx 32 years standing on a forest track, Bai Hokou, Dzanga Sangha Special Dense Forest Reserve...

Anup Shah / Western lowland gorilla (Gorilla gorilla gorilla) dominant male silverback 'Makumba' aged 32 years feeding on sedge grasses in Bai Hokou, Dzanga Sangha Special Dense Forest Reserve, Central African Re...

Anup Shah / Western lowland gorilla (Gorilla gorilla gorilla) sub-adult male 'Kunga' aged 13 years looking at his hand, Bai Hokou, Dzanga Sangha Special Dense Forest Reserve, Central African Republic. December 20...

Anup Shah / Western lowland gorilla (Gorilla gorilla gorilla) sub-adult male 'Kunga' aged 13 years head and shoulders portrait, Bai Hokou, Dzanga Sangha Special Dense Forest Reserve, Central African Republic. Dec...

Anup Shah / Western lowland gorilla (Gorilla gorilla gorilla) juvenile male 'Tembo' aged 4 years reaching into a hole in a tree for rotting wood to feed on, Bai Hokou, Dzanga Sangha Special Dense Forest Reserve,...

Anup Shah / Western lowland gorilla (Gorilla gorilla gorilla) female 'Malui' feeding on rotting wood from a hole in a tree, young gorilla nearby, Bai Hokou, Dzanga Sangha Special Dense Forest Reserve, Central Afr...

Anup Shah / Western lowland gorilla (Gorilla gorilla gorilla) female 'Malui' feeding on rotting wood from a hole in a tree, Bai Hokou, Dzanga Sangha Special Dense Forest Reserve, Central African Republic. Decembe...

Anup Shah / Western lowland gorilla (Gorilla gorilla gorilla) dominant male silverback 'Makumba' aged 32 years feeding on rotten wood from a tree while his juvenile son 'Tembo' aged 4 years waits his turn, Bai Ho...

Anup Shah / Western lowland gorilla (Gorilla gorilla gorilla) juvenile male 'Mobangi' aged 5 years feeding on rotting wood, Bai Hokou, Dzanga Sangha Special Dense Forest Reserve, Central African Republic. Decembe...

Anup Shah / Western lowland gorilla (Gorilla gorilla gorilla) dominant male silverback 'Makumba' aged 32 years standing portrait, Bai Hokou, Dzanga Sangha Special Dense Forest Reserve, Central African Republic. D...

Anup Shah / Western lowland gorilla (Gorilla gorilla gorilla) juvenile male 'Mobangi' aged 5 years feeding on fruit while holding a piece of rotting wood which he will eat later, Bai Hokou, Dzanga Sangha Special...

Anup Shah / Western lowland gorilla (Gorilla gorilla gorilla) dominant male silverback 'Makumba' aged 32 years sitting in a forest clearing, Bai Hokou, Dzanga Sangha Special Dense Forest Reserve, Central African...

Anup Shah / Western lowland gorilla (Gorilla gorilla gorilla) dominant male silverback 'Makumba' aged 32 years feeding on sedge grasses in Bai Hokou, Dzanga Sangha Special Dense Forest Reserve, Central African Re...

Anup Shah / Western lowland gorilla (Gorilla gorilla gorilla) juvenile male 'Mobangi' aged 5 years feeding on fruit while holding a piece of rotting wood which he will eat later, Bai Hokou, Dzanga Sangha Special...

Anup Shah / Western lowland gorilla (Gorilla gorilla gorilla) juvenile male 'Tembo' aged 4 years feeding on sedge grasses in Bai Hokou, Dzanga Sangha Special Dense Forest Reserve, Central African Republic. Decemb...

Anup Shah / Western lowland gorilla (Gorilla gorilla gorilla) juvenile male 'Tembo' aged 4 years feeding on sedge grasses in Bai Hokou, Dzanga Sangha Special Dense Forest Reserve, Central African Republic. Decemb...

Anup Shah / Western lowland gorilla (Gorilla gorilla gorilla) dominant male silverback 'Makumba' aged 32 years feeding on sedge grasses in Bai Hokou, Dzanga Sangha Special Dense Forest Reserve, Central African Re...

Anup Shah / Western lowland gorilla (Gorilla gorilla gorilla) dominant male silverback 'Makumba' aged 32 years sitting in a forest clearing, Bai Hokou, Dzanga Sangha Special Dense Forest Reserve, Central African...

Anup Shah / Western lowland gorilla (Gorilla gorilla gorilla) juvenile male 'Mobangi' aged 5 years sitting on the forest floor feeding on fruit, Bai Hokou, Dzanga Sangha Special Dense Forest Reserve, Central Afri...

Anup Shah / Western lowland gorilla (Gorilla gorilla gorilla) juvenile male 'Tembo' aged 4 years feeding while sitting on the forest floor, Bai Hokou, Dzanga Sangha Special Dense Forest Reserve, Central African R...

Anup Shah / Western lowland gorilla (Gorilla gorilla gorilla) female 'Mopambi' feeding on fruit, Bai Hokou, Dzanga Sangha Special Dense Forest Reserve, Central African Republic. December 2011.

Anup Shah / Western lowland gorilla (Gorilla gorilla gorilla) dominant male silverback 'Makumba' aged 32 years drinking from a river while sub-adult female 'Mosoko' aged 8 years crosses bi-pedally, Bai Hokou, Dza...

Anup Shah / Western lowland gorilla (Gorilla gorilla gorilla) juvenile female 'Bokata' aged 6 years walking bi-pedally to cross a river, Bai Hokou, Dzanga Sangha Special Dense Forest Reserve, Central African Repu...

Anup Shah / Western lowland gorilla (Gorilla gorilla gorilla) dominant male silverback 'Makumba' aged 32 years sitting portrait, Bai Hokou, Dzanga Sangha Special Dense Forest Reserve, Central African Republic. De...

Anup Shah / Western lowland gorilla (Gorilla gorilla gorilla) dominant male silverback 'Makumba' aged 32 years feeding on fruit with his son 'Tembo' aged 4 years sitting in the background, Bai Hokou, Dzanga Sangh...

Anup Shah / Western lowland gorilla (Gorilla gorilla gorilla) juvenile male 'Mobangi' aged 5 years sitting portrait, Bai Hokou, Dzanga Sangha Special Dense Forest Reserve, Central African Republic. December 2011.

Anup Shah / Western lowland gorilla (Gorilla gorilla gorilla) juvenile female 'Bokata' aged 6 years walking bi-pedally to cross a river, Bai Hokou, Dzanga Sangha Special Dense Forest Reserve, Central African Repu...

Anup Shah / Western lowland gorilla (Gorilla gorilla gorilla) juvenile male 'Mobangi' aged 5 years sitting on a fallen tree, Bai Hokou, Dzanga Sangha Special Dense Forest Reserve, Central African Republic. Decemb...

Anup Shah / Western lowland gorilla (Gorilla gorilla gorilla) sub-adult female 'Mosoko' aged 8 years sitting portrait, Bai Hokou, Dzanga Sangha Special Dense Forest Reserve, Central African Republic. December 201...

Anup Shah / Western lowland gorilla (Gorilla gorilla gorilla) juvenile male 'Mobangi' aged 5 years sitting portrait, Bai Hokou, Dzanga Sangha Special Dense Forest Reserve, Central African Republic. December 2011.

Anup Shah / Western lowland gorilla (Gorilla gorilla gorilla) juvenile male 'Mobangi' aged 5 years head and shoulders portrait, Bai Hokou, Dzanga Sangha Special Dense Forest Reserve, Central African Republic. Dec...

Anup Shah / Western lowland gorilla (Gorilla gorilla gorilla) juvenile male 'Mobangi' aged 5 years trying to break into a termite nest, Bai Hokou, Dzanga Sangha Special Dense Forest Reserve, Central African Repub...

Anup Shah / Western lowland gorilla (Gorilla gorilla gorilla) dominant male silverback 'Makumba' aged 32 years sitting in a forest clearing, Bai Hokou, Dzanga Sangha Special Dense Forest Reserve, Central African...

Anup Shah / Western lowland gorilla (Gorilla gorilla gorilla) female 'Malui' sitting in a tree, Bai Hokou, Dzanga Sangha Special Dense Forest Reserve, Central African Republic. December 2011.

Anup Shah / Western lowland gorilla (Gorilla gorilla gorilla) juvenile male 'Mobangi' aged 5 years feeding on sedge grasses in a bai with the rest of his group, Bai Hokou, Dzanga Sangha Special Dense Forest Reser...

Anup Shah / Western lowland gorilla (Gorilla gorilla gorilla) sub-adult female 'Mosoko' aged 8 years sitting on a fallen tree feeding on fruit, Bai Hokou, Dzanga Sangha Special Dense Forest Reserve, Central Afric...

Anup Shah / Western lowland gorilla (Gorilla gorilla gorilla) sub-adult male 'Kunga' aged 13 years feeding on sedge grasses in Bai Hokou, Dzanga Sangha Special Dense Forest Reserve, Central African Republic. Dece...

Anup Shah / Western lowland gorilla (Gorilla gorilla gorilla) juvenile male 'Mobangi' aged 5 years sitting portrait, Bai Hokou, Dzanga Sangha Special Dense Forest Reserve, Central African Republic. December 2011.

Anup Shah / Western lowland gorilla (Gorilla gorilla gorilla) juvenile male 'Tembo' aged 4 years peering through vegetation, Bai Hokou, Dzanga Sangha Special Dense Forest Reserve, Central African Republic. Decemb...

Anup Shah / Western lowland gorilla (Gorilla gorilla gorilla) sub-adult male 'Kunga' aged 13 years standing portrait, Bai Hokou, Dzanga Sangha Special Dense Forest Reserve, Central African Republic. December 2011...

Anup Shah / Western lowland gorilla (Gorilla gorilla gorilla) juvenile male 'Mobangi' aged 5 years feeding on termites, Bai Hokou, Dzanga Sangha Special Dense Forest Reserve, Central African Republic. November 20...

Anup Shah / Western lowland gorilla (Gorilla gorilla gorilla) dominant male silverback 'Makumba' aged 32 years walking bi-pedally across a river, Bai Hokou, Dzanga Sangha Special Dense Forest Reserve, Central Afr...

Anup Shah / Western lowland gorilla (Gorilla gorilla gorilla) dominant male silverback 'Makumba' aged 32 years sitting by a river in forest interior, Bai Hokou, Dzanga Sangha Special Dense Forest Reserve, Central...

Anup Shah / Western lowland gorilla (Gorilla gorilla gorilla) sub-adult male 'Kunga' aged 13 years head and shoulders portrait, Bai Hokou, Dzanga Sangha Special Dense Forest Reserve, Central African Republic. Dec...

Anup Shah / Western lowland gorilla (Gorilla gorilla gorilla) sub-adult female 'Mosoko' aged 8 years sitting on a fallen tree feeding on fruit, Bai Hokou, Dzanga Sangha Special Dense Forest Reserve, Central Afric...