Anup Shah / Western lowland gorilla (Gorilla gorilla gorilla) sub-adult female 'Mosoko' aged 8 years crossing a river, Bai Hokou, Dzanga Sangha Special Dense Forest Reserve, Central African Republic

Dave Watts / Wedge-tailed eagle {Aquila audax fleayi} tasmanian sub species, adult portrait, Australia

Anup Shah / Eastern Chimpanzee (Pan troglodytes schweinfurthii) fourteen year old sub-adult male, named Fundi, making submissive facial gesture, Gombe National Park, Tanzania

Nick Garbutt / Puma (Puma concolor puma) young male standing with mouth open. Estancia Amarga, near Torres del Paine National Park, Patagonia, Chile. December.

Tony Wu / Bowhead whale (Balaena mysticetus), group socialising in shallow water, aerial view. Part of larger group of more than 50 whales congregating, Sea of Okhotsk sub-population, Russia.

Andy Rouse / Bengal tiger (Panthera tigris) male, sub-adult aged three years, face portrait. Ranthambhore National Park, India.

Yashpal Rathore / Bengal tiger (Panthera tigris tigris) sub adult tigress resting at dusk. Kanha National Park, Central India. Camera trap image.

Yashpal Rathore / Bengal tiger (Panthera tigris tigris) sub adult tigress resting at dusk. Kanha National Park, Central India. Camera trap image.

SCOTLAND: The Big Picture / Mountain hare (Lepus timidus), sub-adult in summer coat on moorland, Scotland, UK, August.

Franco Banfi / Striped marlin (Kajikia audax) hunting Sardine (Sardinops sagax) in bait ball. Magdalena Bay, Baja California Sur, Mexico.

Franco Banfi / California sealion (Zalophus californianus) swimming. Magdalena Bay, Baja California Sur, Pacific Ocean, Mexico

Franco Banfi / Striped marlin (Tetrapturus audax) feeding on Sardine (Sardinops sagax). Magdalena Bay, Baja California Sur, Pacific Ocean, Mexico.

Ingo Arndt / Mountain Lion (Puma concolor) sub-adult male, Torres del Paine National Park, Patagonia, Chile

Ingo Arndt / Mountain Lion (Puma concolor) sub-adult feeding on Guanaco (Lama guanicoe) prey, Torres del Paine National Park, Patagonia, Chile. Siena International Photo Awards 2022 - Remarkable Artwork - Storyboa...

Ingo Arndt / Mountain Lion (Puma concolor) sub-adult sisters, Torres del Paine National Park, Patagonia, Chile

Sebastian Kennerknecht / Leopard (Panthera pardus) female feeding on sub-adult male Impala (Aepyceros melampus) prey in tree, Greater Makalali Private Game Reserve, South Africa

Alan Murphy/ BIA / Northern Caracara (Caracara cheriway) chasing sub-adult with prey, Texas

Staffan Widstrand / Wild Wonders of China / Red panda or Lesser panda (Ailurus fulgens) in the humid montane mixed forest, Laba He National Nature Reserve, Sichuan, China

Richard Robinson / Snares crested penguin (Eudyptes robustus) on rock washed with sea surf, Snares (Tine Heke) Sub Antarctic Island Group, Southern Ocean, New Zealand. December 2016. Editorial use only.

Frederique Olivier / Pair of King penguins (Aptenodytes patagonicus) regurgitating and calling with chick nearby, Macquarie Island, Sub-Antarctic Australia.

Frederique Olivier / Two King penguins (Aptenodytes patagonicus) walking along the shore, shot tilts up from feet to head, Macquarie Island, Sub-Antarctic Australia.

Frederique Olivier / Close-up of a Light-mantled sooty albatross (Phoebetria palpebrata) building its nest, Macquarie Island, Sub-Antarctic Australia.

Frederique Olivier / Close-up of a dominant male Southern elephant seal (Mirounga leonina) calling, Macquarie Island, Sub-Antarctic Australia.

Tui De Roy / Snares Crested penguin (Eudyptes robustus) group walking up granite slope into Oleander forest, Station Point colony. Snares Island, New Zealand Sub-Antarctic Islands.

Ingo Arndt / Wind storm in Fortuna Bay area, South Georgia, February 2011

Ingo Arndt / Fisheries Protection Vessel (Pharos) arrived at King Edward Point with King penguins on shoreline, South Georgia Heritage Trust Rat Eradication Project, South Georgia, February 2011

Yva Momatiuk & John Eastcott / Ostrich (Struthio camelus) sub-adults at waterhole in dry season, Etosha National Park, Namibia

Hiroya Minakuchi / King Penguin (Aptenodytes patagonicus) colony, South Georgia Island

Gerry Ellis / Orangutan (Pongo pygmaeus) sub-adult male in trees in haze caused by forest clearing, West Kalimantan, Borneo, Indonesia. October, 2015

Ole Jorgen Liodden / Campbell albatross (Thalassarche impavida) in flight, silhouetted at sunset, near Campbell Islands, Sub-Antarctic New Zealand. February.

Ole Jorgen Liodden / Erect-crested penguin (Eudyptes sclateri) surfacing near Bounty Island, Sub-Antarctic New Zealand, March. Endangered species.

Ole Jorgen Liodden / Campbell albatross (Thalassarche impavida) in flight, silhouetted at sunset, near Campbell Islands, Sub-Antarctic New Zealand. February.

Ole Jorgen Liodden / Light-mantled sooty albatross (Phoebetria palpebrata) in flight, Auckland Island, Sub-Antarctic New Zealand. February.

Ole Jorgen Liodden / Erect-crested penguin (Eudyptes sclateri) portrait on rocks, Antipodes Island, Sub-Antarctic New Zealand, March. Endangered species.

Greg Oakley / Australian King Parrot (Alisterus scapularis) sub-adult male, Victoria, Australia

Sergey Gorshkov / Leopard (Panthera pardus) sub-adult with Impala (Aepyceros melampus) kill in tree, Sabi-sands Game Reserve, South Africa

Tui De Roy / Erect-crested penguins (Eudyptes sclateri) portrait, Proclamation Island, Bounty Islands, New Zealand Sub-Antarctic Islands. Endemic to Antipodes and Bounty Islands. Endangered species.

Tui De Roy / Erect-crested penguins (Eudyptes sclateri) pair on coast, Proclamation Island, Bounty Islands, New Zealand Sub-Antarctic Islands. Endemic to Antipodes and Bounty Islands. Endangered species.

Tui De Roy / Snares Crested penguin (Eudyptes robustus) bathing in freshwater pool, Snares Island, New Zealand Sub-Antarctic Islands. Endemic and vulnerable species.

Tui De Roy / Snares-crested penguin (Eudyptes robustus) walking along rocks, Snares Island, New Zealand Sub-Antarctic Islands. Endemic and vulnerable species.

Tui De Roy / Erect-crested penguin (Eudyptes sclateri) pair. Antipodes Island, New Zealand Sub-Antarctic Islands. Endemic to Antipodes and Bounty Islands. Endangered species.

Tui De Roy / Erect-crested penguins (Eudyptes sclateri) feeding young chick. Antipodes Island, New Zealand Sub-Antarctic Islands. Endemic to Antipodes and Bounty Islands. Endangered species.

Tui De Roy / Erect-crested penguins (Eudyptes sclateri) pair. Proclamation Island, Bounty Islands, New Zealand Sub-Antarctic Islands. Endemic to Antipodes and Bounty Islands. Endangered species.

Tui De Roy / Erect-crested penguin (Eudyptes sclateri) pair in greeting display. Antipodes Island, New Zealand Sub-Antarctic Islands. Endemic to Antipodes and Bounty Islands. Endangered species.

Graeme Guy / White-backed Vulture (Gyps africanus) and sub-adult, Masai Mara, Kenya

Roland Seitre / Young Lion (Panthera leo) in tree, Fathala Reserve, Senegal. Captive, occurs in sub-Saharan Africa, Vulnerable species.

Ammonite / African lioness (Panthera leo) growling and swatting at a sub-adult male approaching a kill, footage taken at night using thermal camera technology, Masai Mara, Kenya.

Ammonite / Group of sub-adult African lions (Panthera leo) chasing and jumping on a mother Hippopotamus (Hippopotamus amphibius) and her calf, footage taken at night using infrared camera technology, Masai Mara,...

Ammonite / Sub-adult African lioness (Panthera leo) chasing and jumping onto the back of a Hippopotamus (Hippopotamus amphibius), footage taken at night using infrared camera technology, Masai Mara, Kenya.

Ammonite / Sub-adult African lioness (Panthera leo) watching, chasing and missing a small group of Warthogs (Phacochoerus aethiopicus), Masai Mara, Kenya.

Mark Hamblin / 2020VISION / Two sub-adult Wild Domestic goats (Capra hircus) play fighting on hillside, Cairngorms National Park, Scotland, UK, February.

Roland Seitre / Dwarf Gibbon (Hylobates klossii) sub-adult in tree, native to Asia

Enrique Lopez-Tapia / The forests of the Modi Khola river valley shrouded in fog. Annapurna Sanctuary, central Nepal, November 2011.

Enrique Lopez-Tapia / Tourists riding domesticated Indian Elephant (Elephas maximas) on safari, Royal Chitwan National Park, Nepal. October 2011.

Anup Shah / Lion-tailed macaque (Macaca silenus) sub-adult male sitting in a tree. Anamalai Tiger Reserve, Western Ghats, Tamil Nadu, India.

Andrew Parkinson / Gannet (Morus bassanus) sub-adults roosting on a coastal rock Shetland Islands, Scotland, UK, July

Anup Shah / Celebes / Black crested macaque (Macaca nigra) sub-adult male yawning, Tangkoko National Park, Sulawesi, Indonesia.

Fiona Rogers / Celebes / Black crested macaque (Macaca nigra) sub-adult male head portrait, Tangkoko National Park, Sulawesi, Indonesia.

Fiona Rogers / Celebes / Black crested macaque (Macaca nigra) sub-adult male head portrait, Tangkoko National Park, Sulawesi, Indonesia.

Anup Shah / Celebes / Black crested macaque (Macaca nigra) sub-adult male sitting on a beach, Tangkoko National Park, Sulawesi, Indonesia.

Anup Shah / Celebes / Black crested macaque (Macaca nigra) sub-adult male resting on a fallen tree, Tangkoko National Park, Sulawesi, Indonesia.

Anup Shah / Celebes / Black crested macaque (Macaca nigra) sub-adult male trying to break into bamboo to feed on ant eggs from a nest, Tangkoko National Park, Sulawesi, Indonesia.

Anup Shah / Celebes / Black crested macaque (Macaca nigra) sub-adult male head and shoulders portrait, Tangkoko National Park, Sulawesi, Indonesia.

Mark Carwardine / Dwarf minke whale, thought to form a yet-to-be named sub-species of the Common minke whale {Balaenoptera acutorostrata}, Ribbon Reefs, Great Barrier Reef, Queensland, Australia

Mark Carwardine / Rear view of Dwarf minke whale, thought to form a yet-to-be named sub-species of the Common minke whale {Balaenoptera acutorostrata}, Ribbon Reefs, Great Barrier Reef, Queensland, Australia

Mark Carwardine / Tail flippers of Dwarf minke whale, thought to form a yet-to-be named sub-species of the Common minke whale {Balaenoptera acutorostrata}, Ribbon Reefs, Great Barrier Reef, Queensland, Australia

Floris van Breugel / Sub-alpine larches (Latrix sp.) in late autumn in North Cascades National Park, Washington, USA. October.

Ole Jorgen Liodden / Royal Penguin (Eudyptes schlegeli) Macquarie Island, Sub-Antarctic Australia. November.

Ole Jorgen Liodden / King Penguin (Aptenodytes patagonicus) swimming underwater past kelp, Macquarie Island, Sub-Antarctic Australia. November.

Ole Jorgen Liodden / Royal Penguin (Eudyptes schlegeli) fighting, Macquarie Island, Sub-Antarctic Australia. November.

Ole Jorgen Liodden / Royal Penguins (Eudyptes schlegeli) fighting, Macquarie Island, Sub-Antarctic Australia. November.

Ole Jorgen Liodden / Royal Penguin (Eudyptes schlegeli) swimming underwater near Macquarie Island, Sub-Antarctic, Australia. November.

Ole Jorgen Liodden / Royal Penguin (Eudyptes schlegeli) amongst king penguins, Macquarie Island, Sub-Antactcita, Australia. November.

Ole Jorgen Liodden / Male Hooker's / New Zealand sea lion (Phocarctos hookeri) in water, Campbell Island, New Zealand, sub Antarctic Islands. November.

Christophe Courteau / West African giraffe {Giraffa camelopardis peralta} poached by local people. Sahel, Niger.

Christophe Courteau / West African giraffes {Giraffa camelopardis peralta} in millet plantation Sahel, Niger.

Christophe Courteau / West African giraffe {Giraffe camelopardis peralta} in local village. Sahel, Niger.

Christophe Courteau / West African giraffe {Giraffe camelopardis peralta} skin detail.

Christophe Courteau / Pair of West African giraffes {Giraffe camelopardis peralta}. Male on right, female on left.

Christophe Courteau / West African giraffe male feeding {Giraffa camelopardis peralta}. Sahel, Niger.

Christophe Courteau / West African giraffes {Giraffa camelopardalis peralta}. Dallol Bosso Nord, Sahel, Niger.

Christophe Courteau / West African giraffe calf {Giraffa camelopardalis peralta}, Niger.

Christophe Courteau / West African giraffe face {Giraffa camelopardis peralta}. Sahel, Niger.

Anup Shah / Western lowland gorilla (Gorilla gorilla gorilla) sub-adult female 'Mosoko' aged 8 years feeding on fruit, Bai Hokou, Dzanga Sangha Special Dense Forest Reserve, Central African Republic. December 201...

Anup Shah / Western lowland gorilla (Gorilla gorilla gorilla) sub-adult female 'Mosoko' aged 8 years drinking in Bai Hokou, Dzanga Sangha Special Dense Forest Reserve, Central African Republic. November 2011.

Anup Shah / Western lowland gorilla (Gorilla gorilla gorilla) sub-adult male 'Kunga' aged 13 years drinking in Bai Hokou, Dzanga Sangha Special Dense Forest Reserve, Central African Republic. November 2011.

Anup Shah / Western lowland gorilla (Gorilla gorilla gorilla) sub-adult female 'Mosoko' aged 8 years feeding on sedge grasses in Bai Hokou, Dzanga Sangha Special Dense Forest Reserve, Central African Republic. No...

Anup Shah / Western lowland gorilla (Gorilla gorilla gorilla) sub-adult male 'Kunga' aged 13 years head and shoulders portrait, Bai Hokou, Dzanga Sangha Special Dense Forest Reserve, Central African Republic. Dec...

Anup Shah / Western lowland gorilla (Gorilla gorilla gorilla) sub-adult male 'Kunga' aged 13 years head and shoulders portrait, Bai Hokou, Dzanga Sangha Special Dense Forest Reserve, Central African Republic. Dec...

Anup Shah / Western lowland gorilla (Gorilla gorilla gorilla) sub-adult male 'Kunga' aged 13 years looking at his hand, Bai Hokou, Dzanga Sangha Special Dense Forest Reserve, Central African Republic. December 20...

Anup Shah / Western lowland gorilla (Gorilla gorilla gorilla) sub-adult male 'Kunga' aged 13 years head and shoulders portrait, Bai Hokou, Dzanga Sangha Special Dense Forest Reserve, Central African Republic. Dec...

Anup Shah / Western lowland gorilla (Gorilla gorilla gorilla) sub-adult female 'Mosoko' aged 8 years feeding on sedge grasses while juvenile male 'Mobangi' aged 5 years feeds on rotten wood in the background, Bai...

Anup Shah / Western lowland gorilla (Gorilla gorilla gorilla) sub-adult female 'Mosoko' aged 8 years feeding on rotting wood, Bai Hokou, Dzanga Sangha Special Dense Forest Reserve, Central African Republic. Decem...

Anup Shah / Western lowland gorilla (Gorilla gorilla gorilla) dominant male silverback 'Makumba' aged 32 years drinking from a river while sub-adult female 'Mosoko' aged 8 years crosses bi-pedally, Bai Hokou, Dza...

Anup Shah / Western lowland gorilla (Gorilla gorilla gorilla) dominant male silverback 'Makumba' aged 32 years drinking from a river watched by sub-adult female 'Mosoko' aged 8 years and juvenile male 'Mobangi' a...

Anup Shah / Western lowland gorilla (Gorilla gorilla gorilla) dominant male silverback 'Makumba' aged 32 years approaching a river while sub-adult female 'Mosoko' aged 8 years and juvenile male 'Mobangi' aged 5 y...