Sven Zacek / Fritillary butterfly (Melitaea sp.) – either Heath or False Heath – at rest on a grass stem. Estonia, Europe, June.

David Kjaer / Pearl-bordered Fritillary butterfly (Boloria euphrosyne) at rest on Bracken with wings open, Bentley Wood, Wiltshire, England, UK

David Kjaer / Pearl-bordered Fritillary butterfly (Boloria euphrosyne) at rest on Bracken with wings closed, Bentley Wood, Wiltshire, England, UK

Paul Hobson / Pearl-bordered fritillary (Boloria euphrosyne) butterfly basking on Bracken (Pteridium aquilinum) frond. North Wales, UK. June.

Paul Hobson / Pearl-bordered fritillary (Boloria euphrosyne) resting on Bracken (Pteridium aquilinum) frond. North Wales, UK. June.

Jussi Murtosaari / Pearl-bordered fritillary (Boloria euphrosyne) feeding from Wild Rosemary (Rhododendron tomentosum), Finland, June.

Andy Sands / Pearl-bordered fritillary (Boloria euphrosyne) roosting on Bracken frond, Hampshire, England, UK, May

Andy Sands / Pearl-bordered Fritillary (Boloria euphrosyne) Roosting on Bluebell flower, Hampshire, England, UK, May

Jussi Murtosaari / Pearl Bordered Fritillary butterfly (Clossiana euphrosyne) male, central Finland, May.

Jussi Murtosaari / Pearl Bordered Fritillary (Clossiana euphrosyne) eating on a flower, central Finland, June.

Jussi Murtosaari / Pearl-bordered Fritillary butterfly (Boloria euphrosyne) female sunbathing in habitat, central Finland, June.

Danny Green / Pearl-bordered fritillary (Boloria selene) sunning on a fern frond, Glenn Afric, Scotland, July. Bookplate from Danny Green's 'The Long Journey North'

Loic Poidevin / Pearl bordered fritillary butterfly (Boloria euphrosyne) on Aster flowers, Vendee, West France

Andy Sands / Pearl-bordered Fritillary (Boloria euphrosyne) on bugle flower. West Sussex, England, June.

Andy Sands / Pearl-bordered Fritillary (Boloria euphrosyne) with wings closed on Yellow Archangel (lamiastrum galeoblodon) flower. West Sussex, England, June.

Andy Sands / Pearl-bordered Fritillary (Boloria euphrosyne) with wings closed on Bugle flower. West Sussex, England, June.

Simon Colmer / Caterpillar larva of Pearl-bordered Fritillary butterfly (Boloria euphrosyne) feeding on leaf of Common Dog-violet (Viola riviniana) UK, July

Simon Colmer / Caterpillar larva of Pearl-bordered Fritillary butterfly (Boloria euphrosyne) on Common Dog-violet (Viola riviniana) leaf, UK

Simon Colmer / Caterpillar larva of Pearl-bordered Fritillary butterfly (Boloria euphrosyne) feeding on Common Dog-violet (Viola riviniana) leaf, UK

Simon Colmer / Pearl-bordered Fritillary butterfly (Boloria euphrosyne) caterpillar, on violet leaf, Sussex, UK, June

Simon Colmer / Pearl-bordered Fritillary butterfly (Boloria euphrosyne) caterpillar, on violet leaf, Sussex, UK, June

David Kjaer / Pearl-bordered Fritillary butterfly (Boloria euphrosyne) at rest on Bracken with wings closed, Bentley Wood, Wiltshire, England, UK

David Kjaer / Pearl-bordered Fritillary butterfly (Boloria euphrosyne) at rest on leaf litter, on woodland floor, Bentley Wood, Wiltshire, England, UK.

Stephen Dalton / Pearl bordered fritillary butterfly (Brenthis / Boloria euphrosyne) on Bugle flowers (Ajuga sp) UK

Andy Sands / Pearl bordered fritillary {Boloria euphrosyne} resting on flower. France.

Steve Knell / Pearl bordered fritillary {Boloria euphrosyne} England

Ross Hoddinott / Pearl bordered fritillary {Boloria euphrosyne} Cornwall, UK

Jussi Murtosaari / Pearl-bordered fritillary (Boloria euphrosyne) butterfly, pair, in flight and nectaring. Jyvaskyla, Central Finland. June.

David Woodfall / Pearl bordered fritillary butterfly (Boloria euphrosyne) on Wild garlic flower, Denbighshire, Wales, UK. May.

David Woodfall / Pearl bordered fritillary butterflies (Boloria euphrosyne) pair mating on Bracken. Denbighshire, Wales, UK. May.

David Woodfall / Pearl bordered fritillary (Boloria euphrosyne) pair on Hawkbit. Denbighshire, Wales, UK. May.

David Woodfall / Pearl bordered fritillary butterfly (Boloria euphrosyne) on Hawkbit flower. Denbighshire, Wales, UK. May.

David Woodfall / Pearl bordered fritillary butterfly (Boloria euphrosyne) on Wild garlic flower, Denbighshire, Wales, UK. May.

David Woodfall / Pearl-bordered fritillary butterfly (Boloria euphrosyne) feeding on Wild garlic (Allium ursinum) flowers. Threatened species. Dark form. Wales, UK. May.

David Woodfall / Pearl-bordered fritillary butterfly (Boloria euphrosyne) feeding on Wild garlic (Allium ursinum) flowers. Wales, UK. May.

David Woodfall / Pearl-bordered fritillary butterfly (Boloria euphrosyne) feeding on flowers. Threatened species, pale form, Wales, UK.May.

David Woodfall / Pearl-bordered fritillary butterfly (Boloria euphrosyne) feeding on Wild garlic (Allium ursinum) flowers. Threatened species. Dark form, Wales, UK. May.

Jussi Murtosaari / Pearl-bordered fritillary butterfly (Boloria euphrosyne), male, Finland, July.