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Sockeye Salmon (Oncorhynchus nerka) group migrating upstream, Adams River, Roderick Haig-Brown Provincial Park, British Columbia, Canada

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Yva Momatiuk & John Eastcott / Sockeye Salmon (Oncorhynchus nerka) group migrating upstream, Adams River, Roderick Haig-Brown Provincial Park, British Columbia, Canada

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Loic Poidevin / Grizzly bear (Ursus arctos horribilis) mother watching cub catching salmon, Katmai National Park, Alaska, USA, August

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Loic Poidevin / Grizzly bear (Ursus arctos horribilis) cubs with salmon, Katmai National Park, Alaska, August. Small reproduction only.

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Loic Poidevin / Grizzly bear (Ursus arctos horribilis) cubs with salmon, Katmai National Park, Alaska, August. Small reproduction only.

Sockeye Salmon (Oncorhynchus nerka) group swimming upstream during spawning run, Adams River, Roderick Haig-Brown Provincial Park, British Columbia, Canada

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Ingo Arndt / Sockeye Salmon (Oncorhynchus nerka) group swimming upstream during spawning run, Adams River, Roderick Haig-Brown Provincial Park, British Columbia, Canada

Sockeye Salmon (Oncorhynchus nerka) migrating upstream, Adams River, Roderick Haig-Brown Provincial Park, British Columbia, Canada

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Yva Momatiuk & John Eastcott / Sockeye Salmon (Oncorhynchus nerka) migrating upstream, Adams River, Roderick Haig-Brown Provincial Park, British Columbia, Canada

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Michel Roggo / Brown bear (Ursus arctos beringianus) fishing for Sockeye salmon in the Ozernaya River, Kamtchatka, Russia, August

A female (in front) and male Sockeye salmon (Oncorhynchus nerka) over eggs in their spawning river. Adams River, British Columbia, Canada, October.

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Alex Mustard,Alex Mustard / A female (in front) and male Sockeye salmon (Oncorhynchus nerka) over eggs in their spawning river. Adams River, British Columbia, Canada, October.

A split level photo of group of Sockeye salmon (Oncorhynchus nerka) fighting their way upstream as they migrate back to the river of their birth to spawn, trees showing autumnal colours. Adams River,...

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Alex Mustard / A split level photo of group of Sockeye salmon (Oncorhynchus nerka) fighting their way upstream as they migrate back to the river of their birth to spawn, trees showing autumnal colours. Adams River,...

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Matthias Breiter / Sockeye Salmon (Oncorhynchus nerka) group jumping up waterfall, Brooks Falls, Katmai National Park, Alaska

Sockeye / Red Salmon (Oncorhynchus nerka) on spawning migration; males surrounding a female. Adams River, British Columbia, Canada, October.

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Michel Roggo / Sockeye / Red Salmon (Oncorhynchus nerka) on spawning migration; males surrounding a female. Adams River, British Columbia, Canada, October.

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Michel Roggo / Sockeye / Red Salmon (Oncorhynchus nerka) on spawning migration. Adams River, British Columbia, Canada, October.

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Michel Roggo / Sockeye / Red Salmon (Oncorhynchus nerka) on spawning migration. Adams River, British Columbia, Canada, October.

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Michel Roggo / Sockeye / Red Salmon (Oncorhynchus nerka) on spawning migration. Adams River, British Columbia, Canada, October.

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Michel Roggo / Sockeye / Red Salmon (Oncorhynchus nerka) on spawning migration. Adams River, British Columbia, Canada, October.

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Donald M. Jones / Grizzly Bear (Ursus arctos horribilis) eating Sockeye Salmon (Oncorhynchus nerka), Brooks Falls, Alaska

Sockeye salmon (Oncorhynchus nerka) split level view of salmon on river bed, annual spawning run, Adams River, British Columbia, Canada, October

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Todd Mintz / Sockeye salmon (Oncorhynchus nerka) split level view of salmon on river bed, annual spawning run, Adams River, British Columbia, Canada, October

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Todd Mintz / Sockeye salmon (Oncorhynchus nerka) split level view of annual spawning run, Adams River, British Columbia, Canada, October

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Todd Mintz / Sockeye salmon (Oncorhynchus nerka) split level view of annual spawning run, Adams River, British Columbia, Canada, October

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Sergey Gorshkov / Sockeye Salmon (Salmonidae) spawning in Lake Kuril, Kamchatka, Far East Russia, August

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Sergey Gorshkov / Sockeye salmon (Oncorhynchus nerka) spawning, Lake Kuril, Kamchatka, Far East Russia, July

Sockeye Salmon (Oncorhynchus nerka) group swimming in fast current during spawning run, Adams River, Roderick Haig-Brown Provincial Park, British Columbia, Canada

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Yva Momatiuk & John Eastcott / Sockeye Salmon (Oncorhynchus nerka) group swimming in fast current during spawning run, Adams River, Roderick Haig-Brown Provincial Park, British Columbia, Canada

Sockeye Salmon (Oncorhynchus nerka) group swimming upstream between forested banks during spawning run, Adams River, Roderick Haig-Brown Provincial Park, British Columbia, Canada

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Yva Momatiuk & John Eastcott / Sockeye Salmon (Oncorhynchus nerka) group swimming upstream between forested banks during spawning run, Adams River, Roderick Haig-Brown Provincial Park, British Columbia, Canada

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Sockeye Salmon (Oncorhynchus nerka) group swimming close to surface during spawning run, Adams River, Roderick Haig-Brown Provincial Park, British Columbia, Canada

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Yva Momatiuk & John Eastcott / Sockeye Salmon (Oncorhynchus nerka) group swimming close to surface during spawning run, Adams River, Roderick Haig-Brown Provincial Park, British Columbia, Canada

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Floris van Breugel / Kokanee Salmon (Oncorhynchus nerka) on Autumn migration up Taylor Creek from Lake Tahoe, California, USA.

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Michel Roggo / Sockeye / Red Salmon (Oncorhynchus nerka) migrating upstream to spawn, Adams river, British Columbia, Canada 2006

Sockeye / Red Salmon (Oncorhynchus nerka) waiting in lake to enter the Adams river for migration upstream, British Columbia, Canada 2006

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Michel Roggo / Sockeye / Red Salmon (Oncorhynchus nerka) waiting in lake to enter the Adams river for migration upstream, British Columbia, Canada 2006

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Michel Roggo / Sockeye / Red Salmon (Oncorhynchus nerka), female digging to lay eggs on spawning ground, Adams river, British Columbia, Canada 2006

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Michel Roggo / Sockeye / Red Salmon (Oncorhynchus nerka), males at spawning ground, Adams river, British Columbia, Canada 2006

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Michel Roggo / Sockeye / Red Salmon (Oncorhynchus nerka) pair spawning on gravel ground, Adams river, British Columbia, Canada, 1990

Sockeye / Red salmon (Oncorhynchus nerka) in resting pool on migration upstream for spawning, Adams river, British Columbia, Canada, 1990

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Michel Roggo / Sockeye / Red salmon (Oncorhynchus nerka) in resting pool on migration upstream for spawning, Adams river, British Columbia, Canada, 1990

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Michel Roggo / Sockeye / Red Salmon (Oncorhynchus nerka) mass migration upstream for spawning, Kenai Peninsula, Alaska, 1991

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Igor Shpilenok / Steller's sea eagle {Haliaeetus pelagicus} about to pick up Sockeye salmon prey in its claws, Kuril Lake, Kamchatka, Far East Russia

Kamchatka brown bear (Ursus arctos beringianus) feeding on Sockeye salmon prey, Kronotsky Nature Reserve, Kamchatka, Far East Russia

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Igor Shpilenok / Kamchatka brown bear (Ursus arctos beringianus) feeding on Sockeye salmon prey, Kronotsky Nature Reserve, Kamchatka, Far East Russia

Steller's sea eagle {Haliaeetus pelagicus} adult and juvenile squabbling over Sockeye salmon prey, Kuril Lake, Kamchatka, Far East Russia

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Igor Shpilenok / Steller's sea eagle {Haliaeetus pelagicus} adult and juvenile squabbling over Sockeye salmon prey, Kuril Lake, Kamchatka, Far East Russia

Kamchatka brown bear (Ursus arctos beringianus) catching Sockeye salmon prey in water, Kronotsky Nature Reserve, Kamchatka, Far East Russia

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Igor Shpilenok / Kamchatka brown bear (Ursus arctos beringianus) catching Sockeye salmon prey in water, Kronotsky Nature Reserve, Kamchatka, Far East Russia

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Igor Shpilenok / Steller's sea eagle {Haliaeetus pelagicus} in flight carrying Sockeye salmon in claws, Kuril Lake, Kamchatka, Far East Russia

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Matthias Breiter / Sockeye Salmon (Oncorhynchus nerka) jumping upstream during spawning season, Brooks Falls, Katmai National Park, Alaska

Grizzly Bear (Ursus arctos horribilis) catching spawning Sockeye Salmon (Oncorhynchus nerka), Brooks River Falls, Katmai National Park, Alaska

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Thomas Mangelsen / Grizzly Bear (Ursus arctos horribilis) catching spawning Sockeye Salmon (Oncorhynchus nerka), Brooks River Falls, Katmai National Park, Alaska

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Matthias Breiter / Sockeye Salmon (Oncorhynchus nerka) jumping upstream during spawning season, Brooks Falls, Katmai National Park, Alaska

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Konrad Wothe / Sockeye Salmon (Oncorhynchus nerka) leaping up Brooks River Falls during spawning season, Katmai National Park, Alaska

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Doc White / Sockeye salmon (Oncorhynchus nerka) three swimming above river bed to spawn. Southeast Alaska, USA. September.

Brown bear (Ursus arctos), fishing for Sockeye salmon (Oncorhynchus nerka) in the Ozernaya River, Kuril Lake, South Kamtchatka Sanctuary, Far East Russia. August.

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Michel Roggo / Brown bear (Ursus arctos), fishing for Sockeye salmon (Oncorhynchus nerka) in the Ozernaya River, Kuril Lake, South Kamtchatka Sanctuary, Far East Russia. August.

Underwater view of Brown bear (Ursus arctos) paw fishing for Sockeye salmon (Oncorhynchus nerka) with paw outstretched. Ozernaya River, Kuril Lake, South Kamtchatka Sanctuary, Far East Russia. August.

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Michel Roggo / Underwater view of Brown bear (Ursus arctos) paw fishing for Sockeye salmon (Oncorhynchus nerka) with paw outstretched. Ozernaya River, Kuril Lake, South Kamtchatka Sanctuary, Far East Russia. August.

Underwater view of Brown bear (Ursus arctos), fishing for Sockeye salmon in the Ozernaya River, Kuril Lake, South Kamtchatka Sanctuary, Russia

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Michel Roggo / Underwater view of Brown bear (Ursus arctos), fishing for Sockeye salmon in the Ozernaya River, Kuril Lake, South Kamtchatka Sanctuary, Russia

Mass of dead Sockeye Salmon (Oncorhynchus nerka) after spawning, British Columbia, Canada, October. Photographed for the Freshwater Project.

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Michel Roggo / Mass of dead Sockeye Salmon (Oncorhynchus nerka) after spawning, British Columbia, Canada, October. Photographed for the Freshwater Project.

Sockeye Salmon (Oncorhynchus nerka) on spawning ground in groundwater channel, with trees cut by beavers Adams river, British Columbia, Canada, October. Photographed for the Freshwater Project.

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Michel Roggo / Sockeye Salmon (Oncorhynchus nerka) on spawning ground in groundwater channel, with trees cut by beavers Adams river, British Columbia, Canada, October. Photographed for the Freshwater Project.

Sockeye Salmon (Oncorhynchus nerka) swimming upstream during spawning run (aerial photo), Adams River, Roderick Haig-Brown Provincial Park, British Columbia, Canada October.

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Ingo Arndt / Sockeye Salmon (Oncorhynchus nerka) swimming upstream during spawning run (aerial photo), Adams River, Roderick Haig-Brown Provincial Park, British Columbia, Canada October.

Fishery workers counting Sockeye Salmon (Oncorhynchus nerka) aerial photo. Adams River, Roderick Haig-Brown Provincial Park, British Columbia, Canada October.

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Ingo Arndt / Fishery workers counting Sockeye Salmon (Oncorhynchus nerka) aerial photo. Adams River, Roderick Haig-Brown Provincial Park, British Columbia, Canada October.

Sockeye Salmon (Oncorhynchus nerka) swimming upstream during spawning run (aerial photo). A few dead animals visible on left side. Adams River, Roderick Haig-Brown Provincial Park, British Columbia, C...

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Ingo Arndt / Sockeye Salmon (Oncorhynchus nerka) swimming upstream during spawning run (aerial photo). A few dead animals visible on left side. Adams River, Roderick Haig-Brown Provincial Park, British Columbia, C...

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Ingo Arndt / Sockeye salmon (Oncorhynchus nerka) group swimming upstream during spawning run, Adams River, Roderick Haig-Brown Provincial Park, British Columbia, Canada October.

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Ingo Arndt / Sockeye salmon (Oncorhynchus nerka) group swimming upstream during spawning run, Adams River, Roderick Haig-Brown Provincial Park, British Columbia, Canada October.

Wildlife Photographer Ingo Arndt taking underwater pictures of sockeye salmon (Oncorhynchus nerka). Adams River, Roderick Haig-Brown Provincial Park, British Columbia, Canada October.

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Ingo Arndt / Wildlife Photographer Ingo Arndt taking underwater pictures of sockeye salmon (Oncorhynchus nerka). Adams River, Roderick Haig-Brown Provincial Park, British Columbia, Canada October.

Sockeye salmon (Oncorhynchus nerka) dead male after spawning migration. Adams River, Roderick Haig-Brown Provincial Park, British Columbia, Canada October.

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Ingo Arndt / Sockeye salmon (Oncorhynchus nerka) dead male after spawning migration. Adams River, Roderick Haig-Brown Provincial Park, British Columbia, Canada October.

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Ingo Arndt / Wildlife Photographer Ingo Arndt taking underwater pictures of sockeye salmon (Oncorhynchus nerka). Adams River, Roderick Haig-Brown Provincial Park, British Columbia, Canada October.

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Ingo Arndt / Sockeye salmon (Oncorhynchus nerka) dead male after spawning migration. Adams River, Roderick Haig-Brown Provincial Park, British Columbia, Canada October.

Two male Grizzly bears (Ursus arctos horribilis) in a river, one trying to steal the salmon catch from the other, Brooks River, Katmai National Park, Alaska, USA, July.

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Eric Baccega / Two male Grizzly bears (Ursus arctos horribilis) in a river, one trying to steal the salmon catch from the other, Brooks River, Katmai National Park, Alaska, USA, July.

Three male grizzly bears (Ursus arctos horribilis) fishing for Sockeye salmon ((Oncorhynchus nerka) leaping up a waterfall in Brooks River, Katmai National Park, Alaska, USA, July.

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Eric Baccega / Three male grizzly bears (Ursus arctos horribilis) fishing for Sockeye salmon ((Oncorhynchus nerka) leaping up a waterfall in Brooks River, Katmai National Park, Alaska, USA, July.

Male Grizzly bear (Ursus arctos horribilis) fishing for Sockeye salmon (Oncorhynchus nerka) leaping up a waterfall in Brooks River, Katmai National Park, Alaska, USA, July.

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Eric Baccega / Male Grizzly bear (Ursus arctos horribilis) fishing for Sockeye salmon (Oncorhynchus nerka) leaping up a waterfall in Brooks River, Katmai National Park, Alaska, USA, July.

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Eric Baccega / Male Grizzly bear (Ursus arctos horribilis) fishing for Sockeye salmon (Oncorhynchus nerka) leaping up a waterfall in Brooks River, Katmai National Park, Alaska, USA, July.

Brown bears (Ursus arctos) fishing for Sockeye salmon (Oncorhynchus nerka) in the Ozernaya River, Kuril Lake, South Kamchatka Sanctuary, Russia, August

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Michel Roggo / Brown bears (Ursus arctos) fishing for Sockeye salmon (Oncorhynchus nerka) in the Ozernaya River, Kuril Lake, South Kamchatka Sanctuary, Russia, August

Brown bear (Ursus arctos) portrait, whilst fishing for sockeye salmon in the Ozernaya River, Kuril Lake, South Kamchatka Sanctuary, Russia, August

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Michel Roggo / Brown bear (Ursus arctos) portrait, whilst fishing for sockeye salmon in the Ozernaya River, Kuril Lake, South Kamchatka Sanctuary, Russia, August

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Michel Roggo / Brown bear (Ursus arctos) fishing for sockeye salmon in the Ozernaya River, Kuril Lake, South Kamchatka Sanctuary, Russia, August

Brown bear (Ursus arctos) with cubs in water, fishing for sockeye salmon in the Ozernaya River, Kuril Lake, South Kamchatka Sanctuary, Russia, August

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Michel Roggo / Brown bear (Ursus arctos) with cubs in water, fishing for sockeye salmon in the Ozernaya River, Kuril Lake, South Kamchatka Sanctuary, Russia, August

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Michel Roggo / Brown bear (Ursus arctos) with cubs in water, fishing for sockeye salmon in the Ozernaya River, Kuril Lake, South Kamchatka Sanctuary, Russia, August

Brown bear (Ursus arctos) fishing for Sockeye salmon (Oncorhynchus nerka) in the Ozernaya River, Kuril Lake, South Kamchatka Sanctuary, Russia, August

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Michel Roggo / Brown bear (Ursus arctos) fishing for Sockeye salmon (Oncorhynchus nerka) in the Ozernaya River, Kuril Lake, South Kamchatka Sanctuary, Russia, August

Brown bear (Ursus arctos) in river to fish for sockeye salmon in the Ozernaya River, Kuril Lake, South Kamchatka Sanctuary, Russia, August

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Michel Roggo / Brown bear (Ursus arctos) in river to fish for sockeye salmon in the Ozernaya River, Kuril Lake, South Kamchatka Sanctuary, Russia, August

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Michel Roggo / Brown bear (Ursus arctos) fishing for sockeye salmon in the Ozernaya River, Kuril Lake, South Kamchatka Sanctuary, Russia, August

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Michel Roggo / Brown bear (Ursus arctos) fishing for sockeye salmon in the Ozernaya River, Kuril Lake, South Kamchatka Sanctuary, Russia, August

Brown bear (Ursus arctos) with head underwater, fishing for sockeye salmon in the Ozernaya River, Kuril Lake, South Kamchatka Sanctuary, Russia, August

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Michel Roggo / Brown bear (Ursus arctos) with head underwater, fishing for sockeye salmon in the Ozernaya River, Kuril Lake, South Kamchatka Sanctuary, Russia, August

Brown bear (Ursus arctos) standing in river fishing for sockeye salmon in the Ozernaya River, Kuril Lake, South Kamchatka Sanctuary, Russia, August

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Michel Roggo / Brown bear (Ursus arctos) standing in river fishing for sockeye salmon in the Ozernaya River, Kuril Lake, South Kamchatka Sanctuary, Russia, August

Brown bear (Ursus arctos) in river fishing for sockeye salmon in the Ozernaya River, Kuril Lake, South Kamchatka Sanctuary, Russia, August

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Michel Roggo / Brown bear (Ursus arctos) in river fishing for sockeye salmon in the Ozernaya River, Kuril Lake, South Kamchatka Sanctuary, Russia, August

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Michel Roggo / Brown bear (Ursus arctos) fishing for Sockeye salmon (Oncorhynchus nerka) in the Ozernaya River, Kuril Lake, South Kamchatka Sanctuary, Russia, August

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Michel Roggo / Brown bear (Ursus arctos) fishing for sockeye salmon in the Ozernaya River, Kuril Lake, South Kamchatka Sanctuary, Russia

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Michel Roggo / Brown bear (Ursus arctos) fishing for sockeye salmon in the Ozernaya River, Kuril Lake, South Kamchatka Sanctuary, Russia, August

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Michel Roggo / Brown bear (Ursus arctos) fishing for sockeye salmon in the Ozernaya River, Kuril Lake, South Kamchatka Sanctuary, Russia, August

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Michel Roggo / Brown bear (Ursus arctos) fishing for sockeye salmon in the Ozernaya River, Kuril Lake, South Kamchatka Sanctuary, Russia, August

Brown bear (Ursus arctos) head underwater fishing for sockeye salmon in the Ozernaya River, Kuril Lake, South Kamchatka Sanctuary, Russia, August

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Michel Roggo / Brown bear (Ursus arctos) head underwater fishing for sockeye salmon in the Ozernaya River, Kuril Lake, South Kamchatka Sanctuary, Russia, August

Brown bear (Ursus arctos) head underwater while fishing for sockeye salmon in the Ozernaya River, Kuril Lake, South Kamchatka Sanctuary, Russia, August

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Michel Roggo / Brown bear (Ursus arctos) head underwater while fishing for sockeye salmon in the Ozernaya River, Kuril Lake, South Kamchatka Sanctuary, Russia, August

Brown bear (Ursus arctos) with its head underwater whilst fishing for sockeye salmon in the Ozernaya River, Kuril Lake, South Kamchatka Sanctuary, Russia

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Michel Roggo / Brown bear (Ursus arctos) with its head underwater whilst fishing for sockeye salmon in the Ozernaya River, Kuril Lake, South Kamchatka Sanctuary, Russia

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Michel Roggo / Brown bear (Ursus arctos) fishing for Sockeye salmon (Oncorhynchus nerka) in the Ozernaya River, Kuril Lake, South Kamchatka Sanctuary, Russia, August

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Michel Roggo / Brown bear (Ursus arctos) fishing for Sockeye salmon (Oncorhynchus nerka) in the Ozernaya River, Kuril Lake, South Kamchatka Sanctuary, Russia, August

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Michel Roggo / Brown bear (Ursus arctos) fishing for Sockeye salmon (Oncorhynchus nerka) in the Ozernaya River, Kuril Lake, South Kamchatka Sanctuary, Russia, August

Sockeye salmon (Oncorhynchus nerka) on spawning migration in the Ozernaya River, Kuril Lake, South Kamchatka Sanctuary, Russia, August

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Michel Roggo / Sockeye salmon (Oncorhynchus nerka) on spawning migration in the Ozernaya River, Kuril Lake, South Kamchatka Sanctuary, Russia, August

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Michel Roggo / Brown bear (Ursus arctos) fishing for Sockeye salmon (Oncorhynchus nerka) in the Ozernaya River, Kuril Lake, South Kamchatka Sanctuary, Russia, August

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Michel Roggo / Brown bear (Ursus arctos) fishing for Sockeye salmon (Oncorhynchus nerka) in the Ozernaya River, Kuril Lake, South Kamchatka Sanctuary, Russia, August

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Michel Roggo / Brown bear (Ursus arctos) fishing for Sockeye salmon (Oncorhynchus nerka) in the Ozernaya River, Kuril Lake, South Kamchatka Sanctuary, Russia, August

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Michel Roggo / Brown bear (Ursus arctos) fishing for Sockeye salmon (Oncorhynchus nerka) in the Ozernaya River, Kuril Lake, South Kamchatka Sanctuary, Russia, August

Pink salmon (Oncorhynchus gorbuscha) and Sockeye salmon (Oncorhynchus nerka) on spawning migration in the Ozernaya River, Kuril Lake, South Kamchatka Sanctuary, Russia, August

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Michel Roggo / Pink salmon (Oncorhynchus gorbuscha) and Sockeye salmon (Oncorhynchus nerka) on spawning migration in the Ozernaya River, Kuril Lake, South Kamchatka Sanctuary, Russia, August

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Michel Roggo / Sockeye salmon (Oncorhynchus nerka) on spawning migration in the Ozernaya River, Kuril Lake, South Kamchatka Sanctuary, Russia, August

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Michel Roggo / Sockeye salmon (Oncorhynchus nerka) on spawning migration in the Ozernaya River, Kuril Lake, South Kamchatka Sanctuary, Russia, August

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Michel Roggo / Brown bear (Ursus arctos) holding Sockeye salmon caught in Ozernaya River, Kuril Lake, South Kamchatka Sanctuary, Russia, August

Brown bear (Ursus arctos) seen from underwater, fishing for Sockeye salmon, Ozernaya River, Kuril Lake, South Kamchatka Sanctuary, Russia, August

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Michel Roggo / Brown bear (Ursus arctos) seen from underwater, fishing for Sockeye salmon, Ozernaya River, Kuril Lake, South Kamchatka Sanctuary, Russia, August

Brown bear (Ursus arctos) seen from underwater, catching Sockeye salmon in the Ozernaya River, Kuril Lake, South Kamchatka Sanctuary, Russia, August

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Michel Roggo / Brown bear (Ursus arctos) seen from underwater, catching Sockeye salmon in the Ozernaya River, Kuril Lake, South Kamchatka Sanctuary, Russia, August

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Michel Roggo / Brown bear (Ursus arctos) seen from underwater, fishing for Sockeye salmon, Ozernaya River, Kuril Lake, South Kamchatka Sanctuary, Russia, August