Jeff Vanuga / Shiras moose (Alces alces shirasi) bull in rut. Grand Teton National Park, Wyoming, USA, October.

Wim van den Heever / Red-and-green macaw (Ara chloropterus) in flight, Pantanal, Brazil. August.

Cyril Ruoso / Common woolly monkey (Lagothrix lagotricha) reaching forward, Ikamaperou Sanctuary, Amazon, Peru.

Anup Shah / African elephant (Loxodonta africana) walking toward camera, Maasai Mara National Reserve, Kenya.

Roger Powell / Small flock of Brent Geese (Branta bernicla) in flight..Texel Island, The Netherlands, Europe

Espen Bergersen / Red fox (Vulpes vulpes) close up portrait, Lapland, Finland. March.

Ole Jorgen Liodden / Tufted puffin (Fratercula cirrhata) swimming on the sea, Commander Island, Russia, September.

Markus Varesvuo / Black-throated diver (Gavia arctica) on water, Finland, May.

George Sanker / Pronghorn (Antilocapra americana) male, Yellowstone National Park, Wyoming, USA, June.

Markus Varesvuo / Willow Grouse (Lagopus lagopus) in flight over snowy landscape, Utsjoki, Finland April

Markus Varesvuo / Red-crowned Crane (Grus japonensis) in flight, Hokkaido, Japan, February

Angelo Gandolfi / Giant Anteater (Myrmecophaga tridactyla) approaching livestock fence, Pantanal. Brazil