Andy Rouse / Maasai Giraffe (Giraffa camelopardalis tippelskirchi) silhouetted by beams of light from setting sun, Maasai Mara, Kenya, Africa. Winner of the African section of the Natures Best Windland Smith Rice...

Theo Bosboom / Blue hour before sunrise over the dunes of Seilebost beach, Isle of Harris, Hebrides, Scotland. February 2018.

John Chan / Great horned owl (Bubo virginianus) chicks stretching and flapping their wings, Southern California, USA, May.

John Chan / Great horned owl (Bubo virginianus) chick watching for parents to return with food, Southern California, USA, June.

John Chan / Peregrine falcon (Falco peregrinus) plucking Dowitcher (Limnodromus) prey, Southern California, USA.

John Chan / California brown pelicans (Pelecanus occidentalis californicus) roosting on a mudflat, with a flock of Elegant terns (Thalasseus elegans) nearby, Southern California, USA May/2019

Espen Bergersen / Aerial shot of two Killer whales (Orcinus orca) spyhopping, with a snowy landscape in the background, Skjervoy, Troms, Norway, December.

Sven Zacek / Eurasian pygmy owl (Glaucidium passerinum) perched on snow covered branch in evening light. Vorumaa, Southern Estonia. November.

Sven Zacek / Mist over bog with conifers in pre-dawn light. Endla Nature Reserve, Jogevamaa, Central Estonia. June 2016.

Sven Zacek / Ural owl (Strix uralensis) female in flight having taken off from nest, in evening light. Tartumaa, Southern Estonia. May.

Sven Zacek / Ural owl (Strix uralensis) female flying with prey in beak. Flying back to nest in evening light. Tartumaa, Southern Estonia. May.

Sven Zacek / Dandelion (Taraxacum officinale), a field of many seedheads highlighted at sunset. Karula National Park, Valgamaa, Southern Estonia. June 2017.

Sven Zacek / Ural owl (Strix uralensis) perched on Alder (Alnus sp) tree at dusk. Tartumaa, Southern Estonia. April.

Jack Dykinga / Backlit silver cholla cactus (Cylindropuntia echinocarpa) surrounded by California poppies (Eschscholzia californica), beneath the eroded boulders that define the park. Joshua Tree National Park, Moja...

Nayan Khanolkar / Bengal tiger (Panthera tigris tigris) in forest clearing. Camera trap image. Pench National Park, Madhya Pradesh, India. January 2018.

Vincent Grafhorst / Minden / Crater of active volcano at twilight, Erta Ale, Danakil Depression, Ethiopia

Guy Edwardes / Rock formations at Kimmeridge Bay with Clavell Tower in background, Isle of Purbeck, Jurassic Coast, Dorset, England, UK. December 2010.

Guy Edwardes / Rock formations at Kimmeridge Bay, Clavell Tower in background, Isle of Purbeck, Jurassic Coast, Dorset, England, UK. December 2010.

Guy Edwardes / Sea stacks, Mangurstadh, Aird Feinis, Isle of Lewis, Outer Hebrides, Scotland, UK. March 2014.

Guy Edwardes / Eilean Glas Lighthouse, Isle of Scalpay, Outer Hebrides, Scotland, UK. March 2015.

Guy Edwardes / View across Loch Duich to Eilean Donan Castle, Kyle of Lochalsh, Highlands, Scotland, Uk. July 2012.

Guy Edwardes / Coastal cottages at twilight, Portree, Isle of Skye, Inner Hebrides, Scotland, UK. January 2014.

Guy Edwardes / Gaada Stack, Foula, Shetland, Scotland, UK. August, 2016.

Guy Edwardes / Gaada Stack, Foula, Shetland, Scotland, UK. August.

Guy Edwardes / Butt of Lewis lighthouse, Port Nis / Port of Ness, Isle of Lewis, Outer Hebrides, Scotland, UK. March, 2015.

Sergey Gorshkov / Moonrise at twilight, Putoransky State Nature Reserve, Putorana Plateau, Siberia, Russia

Theo Bosboom / Emerald damselfly (Lestes sponsa) in habitat, at twilight, Hondenven, Tubbergen, the Netherlands, August.

Theo Bosboom / Emerald damselfly (Lestes sponsa) in habitat in early morning twilight, Hondenven, Tubbergen, the Netherlands, August.

Theo Bosboom / Banded demoiselles (Calopteryx splendens) males roosting among common nettles, Roosendaal, the Netherlands. July.

Theo Bosboom / Blue damselflies (Coenagrionidae) resting in heather flowers in dawn twilight, Hatertse vennen near Nijmegen, the Netherlands, August.

Michael Hutchinson / European hedgehog (Erinaceus europaeus) in garden at dusk, with house in the background, Bristol, England, UK, 2016.

Theo Bosboom / Limpets (Patellidae) exposed at low tide on the coast of Playa de Ursa, Portugal. Finalist in the Animals in their Environment Category of the Wildlife Photographer of the Year Awards (WPOY) Competiti...

Jack Dykinga / Massive wildflower display with Lanceleaf monolopia 1+Monolopia lanceolata+2, and Orange fiddle neck 1+Amsinckia intermedia+2 The Temblor Range, also carpeted in flowers, in evening light in the backg...

Frederique Olivier / Adelie penguins (Pygoscelis adeliae) walking in front of a blue iceberg, Adelie Land, Antarctica, January.

Philippe Clement / Flock of Jackdaws (Corvus monedula) congregating at dusk at communal roost, Belgium, December.

Sandesh Kadur / Two male King cobras (Ophiophagus hannah) fighting, Agumbe, Karnartaka, India.

John & Mary-Lou Aitchison / Flock of Barnacle geese (Branta leucopsis) flying over the sea, Islay, Inner Hebrides, Scotland, UK, November.

Ingo Arndt / Grizzly bear (Ursus arctos horribilis) silhouette at sunset, Lake Clark National Park, Alaska, USA, July.

Ingo Arndt / Olive ridley sea turtles (Lepidochelys olivacea) females coming ashore at twilight during an arribada (mass nesting event) to lay eggs, Pacific Coast, Ostional, Costa Rica.

Ingo Arndt / Grizzly Bear (Ursus arctos horribilis) waiting for salmon under moonlit sky, Lake Clark National Park, Alaska, USA, August

Ingo Arndt / Grizzly Bear (Ursus arctos horribilis) waiting for salmon at full moon, Lake Clark National Park, Alaska, USA, August

Ingo Arndt / Rising full moon at Lake Clark National Park, Alaska, USA, August 2010

Tim Fitzharris / Arch on beach, El Matador State Beach, California

Espen Bergersen / Humpback whales (Megaptera novaeangliae) bubble net / lunge feeding on herring (Clupea harengus) at dusk, pod co-operative feeding, Kvaloya, Troms, Norway, November

Espen Bergersen / Killer whale / Orca (Orcinus orca) male surfacing at sunset, Kvaloya, Troms, Norway October

David Tipling / Wandering albatross (Diomedea exulans) a pair displaying at dusk, Albatross Island, South Georgia January

David Tipling / Wandering albatross (Diomedea exulans) pair displaying at dusk in ritual bonding behaviour, Albatross Island in Bay of Isles, South Georgia January

Neil Aldridge / Two Impalas (Aepyceros melampus) ewes grooming each other in evening light on South Africa's Kruger National Park.

David Pattyn / Gannet (Morus bassanus) coming in to land in the colony in evening light, Great Saltee, Saltee Islands, County Wexford, Republic of Ireland, June

Jordi Chias / Split level view of Blue shark (Prionace glauca) at surface at sunset, Azores Islands, Portugal, Atlantic Ocean

Floris van Breugel / Sunset on the outer coast of Chichagof Island with the White Sulphur Hotsprings in the distance, South East Alaska, USA, July

Floris van Breugel / Ancient Bristlecone pine (Pinus aristata) clings to life, with a grand view of endless sage valleys, White Mountains, California, USA, July

Danny Green / Atlantic Puffins (Fratercula arctica) resting on a cliff top in the sunset, Sule Skerry, Scotland, UK, July

Danny Green / Atlantic Puffin (Fratercula arctica) preening in late evening light, Heimaey, Westman Isles, Iceland, June

Bence Mate / Group of Little egrets (Egretta garzetta) displaying its ornamental feathers, Lake Csaj, Kiskunsagi National Park, Hungary. February.

Bence Mate / Flock of Great egrets (Ardea alba) at twilight, Lake Csaj, Kiskunsagi National Park, Hungary. May.

Bence Mate / Great egret (Ardea alba) at twilight, Lake Csaj, Kiskunsagi National Park, Hungary. July.

Wim van den Heever / Table Mountain viewed from across the sea at sunset, Cape Town, South Africa

Bence Mate / Black crowned night heron (Nycticorax nycticorax) at twilight with other herons in the background, including Grey heron (Ardea cinerea) in flight, Lake Csaj, Kiskunsagi National Park, Pusztaszer, Hun...

Bence Mate / Grey heron (Ardea cinerea) with lightning storm at night. Lake Csaj, Kiskunsagi National Park, Pusztaszer, Hungary. May 2012.

Bence Mate / Lightning storm over Lake Csaj, Kiskunsagi National Park, Pusztaszer, Hungary. May 2012.

Bence Mate / Bittern (Botaurus stellaris) with Great egret (Ardea alba) in the background, Lake Csaj, Kiskunsagi National Park, Pusztaszer, Hungary.

Bence Mate / Grey heron (Ardea cinerea) face to face in lake, at night with stars, Lake Csaj, Kiskunsagi National Park, Pusztaszer, Hungary. July.

Niall Benvie / Cliff at Vik beach, light by torchlight, Iceland, March 2015.

Niall Benvie / Birch (Betula) at moonrise, Loch Shieldaig, Torridon, Scotland

Niall Benvie / Queen Mary's Tree, a veteran Sweet chestnut (Castanea sativa) over 600 years old, Cumbernauld, Scotland, UK, July.

Espen Bergersen / Gulls (Laridae) seaching for food in water containing blood coming from a large fishing boat, Tromvik, Kvaloya, Troms, Norway, November 2014.

Wim van den Heever / Quiver Tree forest, (Aloe dichotoma) at dusk. Keetmanshoop, Namibia.

Wim van den Heever / Giant anteater (Myrmecophaga tridactyla) walking with its baby on the back at twilight, Pantanal, Brazil.

Wim van den Heever / African elephant (Loxodonta africana) grazing in an open field with a full moon rising near Baobab tree (Adansonia digitata). With tourists on safari under the tree. Savuti Marsh, Botswana, May 2014.

Wim van den Heever / Plains zebra (Equus quagga) group of four including foal at sunset, Savuti Marsh, Botswana.

Wim van den Heever / Plains zebra (Equus quagga) at sunset, Savuti Marsh, Botswana.

Solvin Zankl / English oak tree (Quercus robur) silhouetted against orange sky with star trails, Nauroth, Germany, October.

Brent Stephenson / Sunset off Scott Island, with floating pieces of sea ice, Scott Inlet, Baffin Island, Canadian Arctic, August 2015.

Claudio Contreras / Firewheel (Gaillardia pulchella) flowers at twilight, Laredo Borderlands, Texas, USA. April

Ben Hall / Kirkjufell mountain, landscape at dawn with waterfall in foreground, Snaefellsnes peninsula, Iceland, January 2015

Doug Perrine / Red hot lava from Kilauea Volcano flowing into ocean at West Kailiili, Hawaii Volcanoes National Park, Big Island, Hawaiian Islands, USA.

Jack Dykinga / Lightning storm at twilight with Saguaro cactus (Carnegiea gigantea) Avra Valley, Saguaro National Park, Tucson Mountains, Arizona, USA. June 2015. Long exposure with lightning trigger.

Sven Zacek / Low angle shot of Ural owl (Strix uralensis) female returning to its nest at night. Southern Estonia, May.

Claudio Contreras / Caribbean flamingo (Phoenicopterus ruber) preparing to sleep, Ria Lagartos Biosphere Reserve, Yucatan Peninsula Mexico, July. Bookplate.

Bence Mate / Little egret (Egretta garzetta) and Grey herons (Ardea cinerea) reflected in lake at twilight, Lake Csaj, Pusztaszer, Hungary. June. Winner of the Portfolio category of the Terre Sauvage Nature Images...

Bence Mate / Little egret (Egretta garzetta) shaking water off wings at sunset, Lake Csaj, Pusztaszer, Hungary. May. Winner of the Portfolio category of the Terre Sauvage Nature Images Awards competition 2015.

Colin Varndell / Badger (Meles meles) with face lit up at sunset. Dorset, UK July.

Espen Bergersen / Svalbard reindeer (Rangifer tarandus platyrhynchus) in distance at sunset, Svalbard, Norway. April.

Angelo Gandolfi / White stork (Ciconia ciconia), nesting colony to provide alternative nesting area whilst the building they normally nest on is under repair. Extremadura, Spain, April.

Sven Zacek / Broken ice covered in snow at sunset, on the shore of Lake Vortsjarv, gathered to shore by extreme winds, Estonia. December 2014.

Sven Zacek / Eilean Donan Castle lit up at twilight with dark clouds. Scotland, UK, September 2013.

Sven Zacek / Roe deer (Capreolus capreolus) buck with velvet antlers and fawn at sunset, Tartumaa, March 2013.

Nick Upton / Mute swan (Cygnus olor) female and a young cygnet foraging in sunset light, Kennet and Avon canal, Caen Hill, Wiltshire, UK, June.

Pete Oxford / Three granite peaks of Paine mountain range, Torres del Paine National Park, Patagonia, Chile.

Jack Dykinga / Great horned owl (Bubo virginianus) chicks in nest in Saguaro cactus (Carnegiea gigantea) at sunset, Santa Catalina Mountains, Arizona, USA, May.

Chris & Monique Fallows / Long-beaked common dolphin (Delphinus capensis) school off the coast of False Bay, with dramatic sky, South Africa, May.

Ross Hoddinott / Trevose Lighthouse at dusk, long exposure at high tide, Trevose, Cornwall, UK. January 2015.

Ross Hoddinott / Ice on black volcanic sand beach with waves, Jokulsarlon, Southeast Iceland, February.

Peter Cairns / Twilight over Flakstad beach, Lofoten, Norway. February 2014.