Roger Powell / Ruff (Philomachus pugnax) male showing off his ruff in a courtship display behaviour. Varanger, Norway, June.

Roger Powell / Ruff (Philomachus pugnax) male in full display. This was the dominant male at the lek. Varanger, Norway, June.

Roger Powell / Three male Ruffs (Philomachus pugnax) preparing to display on the communal lekking ground, Varanger plateau, arctic Norway. June

Roger Powell / Ruff (Philomachus pugnax) male displaying to attract the attention of a Reeve (female) nearby, Varanger plateau, Arctic Norway. June

Alan Williams / Ruff (Philomachus pugnax) male foraging along shore in winter plumage, Lancashire, UK, February

Loic Poidevin / Ruff (Philomachus pugnax) males displaying and fighting each other at a lek, Finland. May.

Loic Poidevin / Ruff (Philomachus pugnax), male displaying at lek. Varanger, Norway. May.

Loic Poidevin / Ruff (Philomachus pugnax) male in courtship display at lek. Varanger, Norway. May. Small repro only

Loic Poidevin / Ruff (Philomachus pugnax) male in courtship display at lek. Varanger, Norway. May. Small repro only.

Loic Poidevin / Ruff (Philomachus pugnax) three males in courtship display at lek. Varanger, Norway. May.

Loic Poidevin / Ruff (Philomachus pugnax) male in courtship display at lek. Varanger, Norway. May. Small repro only.

Loic Poidevin / Ruff (Philomachus pugnax) male in courtship display at lek. Varanger, Norway. May.