Yashpal Rathore / Kottigehar dancing frog (Micrixalus kottigeharensis) male calling and displaying to attract female by waving foot, Agumbe, Western Ghats, India.

Philippe Clement / Red deer (Cervus elaphus) stags fighting, Scotland, UK, May.

Tui De Roy / Volcan Alcedo Giant Tortoise (Chelonoidis nigra vandenburghi), Alcedo Volcano, Isabela Island, Galapagos Islands, Ecuador

Guy Edwardes / Snowy owl (Bubo scandiacus) walking on ground in snow, one foot raised, Ontario, Canada, January.

Andrew Parkinson / Mountain hare (Lepus timidus) grooming itself, with back foot raised, Cairngorms National Park, Scotland, UK, February. Highly Commended in the Animal Behaviour category of the British Wildlife Photo...

Tui De Roy / Blue-footed Booby (Sula nebouxii) performing foot-lifting courtship display, Seymour Island, Galapagos Islands, Ecuador

Tui De Roy / Blue-footed Booby (Sula nebouxii) performing foot-lifting courtship display, Seymour Island, Galapagos Islands, Ecuador

Tui De Roy / Blue-footed Booby (Sula nebouxii) feet, Seymour Island, Galapagos Islands, Ecuador

Tui De Roy / Blue-footed Booby (Sula nebouxii) incubating eggs, Santa Cruz Island, Galapagos Islands, Ecuador

Cyril Ruoso / Mute swan (Cygnus olor) feeding underwater, Burgundy, France, March.

Peter Bassett / King penguin (Aptenodytes patagonicus) standing on its heels on a rock to keep its feet warm, Salisbury Plain, South Georgia.

Sue Daly / Close-up of the limbs of a Spiny starfish (Marthasterias glacialis) showing its many tube feet moving over the seabed, Sark, British Channel Islands, UK, August.