Andy Rouse / Asian elephant (Elaphus maximus) older female dominating young calf, captive, Pinnawala Elephant Orphanage, Sri Lanka

Yva Momatiuk & John Eastcott / Guanaco (Lama guanicoe) grazing on grassy slope with dominant male keeping watch of its herd, Torres del Paine National Park, Chile

Theo Allofs / Indian Wild Ass (Equus hemionus khur) dominant male, Indian Wild Ass Sanctuary, Little Rann of Kutch, India

Theo Allofs / Orangutan (Pongo pygmaeus) dominant male sitting on boardwalk during rain, Tanjung Puting National Park, Borneo, Malaysia

Theo Allofs / Bengal Tiger (Panthera tigris tigris) dominant male, resting on dry leaves in bamboo forest, April, dry season, Bandhavgarh National Park, India

Philippe Clement / Koppie (rhyolite hill) dominating the landscape, Kruger NP, South Africa

Philippe Clement / Koppie (rhyolite hill) dominating the landscape, Kruger NP, South Africa

Jim Brandenburg / Arctic Wolf (Canis lupus) alpha male named Buster near den site, Ellesmere Island, Nunavut, Canada

Jim Brandenburg / Arctic Wolf (Canis lupus) alpha male named Buster near den site, Ellesmere Island, Nunavut, Canada