Yukihiro Fukuda / Japanese giant salamander (Andrias japonicus) in river, Japan, March

Yukihiro Fukuda / Japanese giant salamander (Andrias japonicus) reaching up to the surface to breathe, Japan, August

Yukihiro Fukuda / Japanese giant salamander hunting at night in a river, Japan, May

Yukihiro Fukuda / Two male Japanese giant salamanders (Andrias japonicus) fertilizing the eggs of a female, Japan, September

Yukihiro Fukuda / Male Japanese giant salamander (Andria japonicus) emerging from a nest hole in order to breathe, Japan, August

Yukihiro Fukuda / Male Japanese giant salamanders (Andrias japonicus) eating the eggs of another pair, Hino River, Tottori-ken, Japan, September.

Yukihiro Fukuda / Male Japanese giant salamander (Andrias japonicus) protecting its nest from another intruding male, Hino River, Tottori-ken, Japan, September.

Cyril Ruoso / Japanese giant salamander (Andrias japonicus) nest with eggs, Honshu Japan

Cyril Ruoso / Japanese giant salamander (Andrias japonicus) nest with eggs and adult during spawning, Honshu, Japan, September.

Cyril Ruoso / Japanese giant salamander (Andrias japonicus) nest with eggs and adult during spawning, Honshu, Japan, September.

Cyril Ruoso / Japanese giant salamander (Andrias japonicus) surfacing to breathe, Honshu, Japan, August.

Cyril Ruoso / Japanese giant salamander (Andrias japonicus) Honshu, Japan.