Charlie Summers / Arctic grayling (Thymallis arcticus) males during the annual spawning run, North Park, Colorado, USA, June.
01517804
Neil Aldridge / Ring-necked pheasant (Phasianus colchicus) male lies dead on the side of a road near shooting estate, southern England, UK. November. Of the millions of pheasants released into the British countryside...
01514845
Charlie Summers / Arctic grayling (Thymallus arcticus) male with dorsal fin raised, making his way upstream during the annual spawning run. North Park, Colorado creek, USA, June.
01514844
Charlie Summers / Arctic grayling (Thymallus arcticus) male going upstream during the annual spawning run. North Park, Colorado, USA, June.
01514837
Charlie Summers / Male Arctic grayling (Thymalus arcticus) swimming alongside a female during the annual spawning run in a North Park, Colorado, USA, June.
01507897
Nick Upton / Adult male Eurasian beaver (Castor fiber) with identifying ear tag swimming at sunset among felled trees dammed stream within large enclosure, Devon Beaver Project, run by Devon Wildlife Trust, Devon,...
01507896
Nick Upton / Adult male Eurasian beaver (Castor fiber) with identifying ear tag swimming at sunset among felled trees dammed stream within large enclosure, Devon Beaver Project, run by Devon Wildlife Trust, Devon,...
01506412
Nick Upton / Eurasian beaver (Castor fiber) male using its teeth to strip bark from a Willow tree (Salix sp.) in woodland enclosure at night, Devon Beaver Project, run by Devon Wildlife Trust, Devon, UK, April. Ta...
00546107
Donald M. Jones / White-tailed Deer (Odocoileus virginianus) buck running, central Montana
00545899
Donald M. Jones / Rocky Mountain Elk (Cervus canadensis nelsoni) bull running, North America
00545880
Donald M. Jones / Rocky Mountain Elk (Cervus canadensis nelsoni) bull running, North America
00543546
Mark Raycroft / German Shorthaired Pointer (Canis familiaris) running through snow
00543405
Mark Raycroft / Boxer (Canis familiaris) brindle male playing with stick
01484794
Charlie Summers / Arctic grayling (Thymallus arcticus) male in spawning stream during the annual run, North Park, Colorado, USA, June.
01484793
Charlie Summers / Arctic grayling (Thymallus arcticus) male leaping over rocks to reach waterfall during the annual spawning run, North Park, Colorado, USA, June.
01484792
Charlie Summers / Arctic grayling (Thymallus arcticus) male travelling upstream during the annual spawning run, North Park, Colorado, USA, June.
00542274
Theo Allofs / Springbok (Antidorcas marsupialis) male with vegetation entangled in horns, Nxai Pan National Park, Botswana
00538669
Anup Shah / Eastern Chimpanzee (Pan troglodytes schweinfurthii) nineteen year old female, named Imani, with her two year old son, named Ipo, responding to attack by male, Gombe National Park, Tanzania
00538370
Vincent Grafhorst / Hippopotamus (Hippopotamus amphibius) bull running, Chobe National Park, Botswana
00537196
Yva Momatiuk & John Eastcott / Wild Horse (Equus caballus) stallions running together on arid plateau, Pryor Mountain Wild Horse Range, Montana
00537061
Sumio Harada / Bighorn Sheep (Ovis canadensis) rams chasing ewe downhill in snow, Glacier National Park, Montana
00535565
Donald M. Jones / White-tailed Deer (Odocoileus virginianus) buck running across prairie, Montana
00535564
Donald M. Jones / White-tailed Deer (Odocoileus virginianus) buck running through creek, Montana
00535436
Donald M. Jones / Rocky Mountain Elk (Cervus canadensis nelsoni) bull running downriver, Canada
00535410
Donald M. Jones / Bighorn Sheep (Ovis canadensis) rams chasing ewe during rut, Montana
00535683
Duncan Usher / Western Roe Deer (Capreolus capreolus) buck running out of field, Germany
01447196
Jean E. Roche / Man running from Camargue cow in the bull ring, Camargue, France, July 2012.
00531860
Mark Raycroft / White-tailed Deer (Odocoileus virginianus) buck fleeing with tail raised, North America
00531843
Mark Raycroft / White-tailed Deer (Odocoileus virginianus) buck fleeing with tail raised, North America
00531834
Mark Raycroft / Bighorn Sheep (Ovis canadensis) ram running, North America
00531599
Kevin Schafer / Magellanic Penguin (Spheniscus magellanicus) pair courting, Chile
00530764
Michael Quinton / Grizzly Bear (Ursus arctos horribilis) male running after female and yearling cubs, possibly to kill cubs, Alaska
01443494
Wild Wonders of Europe / Widstr / Bull (Bos taurus) in snow, Aurochs breeding site run by The Taurus Foundation, Keent Nature Reserve, The Netherlands.
01443493
Wild Wonders of Europe / Widstr / Bull (Bos taurus) profile in snow, Aurochs breeding site run by The Taurus Foundation, Keent Nature Reserve, The Netherlands.
01443492
Wild Wonders of Europe / Widstr / Bull (Bos taurus) in snow, Aurochs breeding site run by The Taurus Foundation, Keent Nature Reserve, The Netherlands.
01443491
Wild Wonders of Europe / Widstr / Bull (Bos taurus) in snow, Aurochs breeding site run by The Taurus Foundation, Keent Nature Reserve, The Netherlands.
01443490
Wild Wonders of Europe / Widstr / Bull (Bos taurus) in snow, Aurochs breeding site run by The Taurus Foundation, Keent Nature Reserve, The Netherlands.
01443479
Wild Wonders of Europe / Widstr / Bull (Bos taurus) in snow, Aurochs breeding site run by The Taurus Foundation, Keent Nature Reserve, The Netherlands.
01443470
Wild Wonders of Europe / Widstr / Second generation cross-breed bull (Bos taurus), Aurochs breeding site run by The Taurus Foundation, Keent Nature Reserve, The Netherlands.
01443469
Wild Wonders of Europe / Widstr / Second generation cross-breed bull (Bos taurus), Aurochs breeding site run by The Taurus Foundation, Keent Nature Reserve, The Netherlands.
01443468
Wild Wonders of Europe / Widstr / Second generation cross-breed bull (Bos taurus), Aurochs breeding site run by The Taurus Foundation, Keent Nature Reserve, The Netherlands.
01443467
Wild Wonders of Europe / Widstr / Second generation cross-breed bull (Bos taurus), Aurochs breeding site run by The Taurus Foundation, Keent Nature Reserve, The Netherlands.
01443464
Wild Wonders of Europe / Widstr / Second generation cross-breed bull (Bos taurus) profile, Aurochs breeding site run by The Taurus Foundation, Keent Nature Reserve, The Netherlands.
01443459
Wild Wonders of Europe / Widstr / Second generation cross-breed bull (Bos taurus) profile in mist, Aurochs breeding site run by The Taurus Foundation, Keent Nature Reserve, The Netherlands, September.
01443458
Wild Wonders of Europe / Widstr / Second generation cross-breed bull (Bos taurus) profile in mist, Aurochs breeding site run by The Taurus Foundation, Keent Nature Reserve, The Netherlands, September.
01443457
Wild Wonders of Europe / Widstr / Second generation cross-breed bull (Bos taurus) bred for the Aurochs breeding site run by The Taurus Foundation, Keent Nature Reserve, The Netherlands.
01443456
Wild Wonders of Europe / Widstr / Second generation cross-breed bull (Bos taurus) bred for the Aurochs breeding site run by The Taurus Foundation, Keent Nature Reserve, The Netherlands, September.
01443447
Wild Wonders of Europe / Widstr / Second generation crossbreed bull (Bos taurus) seen from wing mirror, bred for the Aurochs breeding site run by The Taurus Foundation, Keent Nature Reserve, The Netherlands.
01443446
Wild Wonders of Europe / Widstr / Second generation cross-breed bull (Bos taurus), Aurochs breeding site run by The Taurus Foundation, Keent Nature Reserve, The Netherlands.
01443445
Wild Wonders of Europe / Widstr / Second generation cross-breed bull (Bos taurus), Aurochs breeding site run by The Taurus Foundation, Keent Nature Reserve, The Netherlands.
01443444
Wild Wonders of Europe / Widstr / Second generation cross-breed bull (Bos taurus), Aurochs breeding site run by The Taurus Foundation, Keent Nature Reserve, The Netherlands.
01445599
Gertrud & Helmut Denzau / Male Goitered gazelle (Gazella subgutturosa) running, Badkhyz Reserve, Turkmenistan. Vulnerable species.
00524772
Pete Oxford / African Lion (Panthera leo) male running, Inkwenkwezi Private Game Reserve, Eastern Cape, South Africa
01433117
Ernie Janes / Sri Lanka Jungle Fowl (Gallus lafayetii) on sandy river bank, Sri Lanka
00521697
Donald M. Jones / White-tailed Deer (Odocoileus virginianus) buck running, northern Rocky Mountains, Canada
00521643
Donald M. Jones / Pronghorn Antelope (Antilocapra americana) male running with grass covered head, western Montana
00455551
Cyril Ruoso / Golden Snub-nosed Monkey (Rhinopithecus roxellana) male displaying, Qinling Mountains, China
00455547
Cyril Ruoso / Golden Snub-nosed Monkey (Rhinopithecus roxellana) male displaying, Qinling Mountains, China
00446760
Cyril Ruoso / Coyote (Canis latrans) male running to calf carcass and scaring off Turkey Vultures (Cathartes aura), southern Texas
00446761
Cyril Ruoso / Coyote (Canis latrans) male running to calf carcass and scaring off Turkey Vultures (Cathartes aura), southern Texas
00621661
Cyril Ruoso / Whooper Swan (Cygnus cygnus) male chasing intruder from nesting area, east Iceland
00620945
Cyril Ruoso / Red Deer (Cervus elaphus) stag running, Denmark
00620064
Cyril Ruoso / Bonobo (Pan paniscus) nine year old male running through vegetation, ABC Sanctuary, Democratic Republic of the Congo
01430618
Jussi Murtosaari / Poplar Admiral (Limenitis populi) male run over by a car, Finland, June
00513072
Richard Du Toit / Ostrich (Struthio camelus) male, Rietvlei Nature Reserve, Gauteng, South Africa
00486460
Mark Raycroft / White-tailed Deer (Odocoileus virginianus) buck running, North America
00486459
Mark Raycroft / White-tailed Deer (Odocoileus virginianus) buck running, North America
00482024
Richard Du Toit / African Elephant (Loxodonta africana) bull chasing Hippopotamus (Hippopotamus amphibius) group, Kruger National Park, South Africa
00482010
Richard Du Toit / Springbok (Antidorcas marsupialis) male herd running, Kalahari, Northern Cape, South Africa
00481683
Colin Monteath / Golden Eagle (Aquila chrysaetos) used for hunting, with Kazak handler competing at winter festival, Mongolia
00481670
Colin Monteath / Red Deer (Cervus elaphus) bulls running up hill in winter, Hustai National Park, Mongolia
00479891
Mark Raycroft / White-tailed Deer (Odocoileus virginianus) buck fleeing with tail raised, North America
00479867
Mark Raycroft / White-tailed Deer (Odocoileus virginianus) buck fleeing with tail raised, North America
00477747
Mark Raycroft / German Shepherd (Canis familiaris) male running in snow
00477716
Mark Raycroft / Dachshund (Canis familiaris) male running
00477715
Mark Raycroft / Dachshund (Canis familiaris) male running
00475950
Pete Oxford / Impala (Aepyceros melampus) male and females running through water, Moremi Game Reserve, Okavango Delta, Botswana. Sequence 2 of 2
00475947
Pete Oxford / Waterbuck (Kobus ellipsiprymnus) male running, Moremi Game Reserve, Okavango Delta, Botswana
00465592
Richard Du Toit / Ostrich (Struthio camelus) males running during dispute, Kalahari, Northern Cape, South Africa
01342041
Mark Taylor / Pack of street dogs following after a bitch on heat in a "bitch run". Coyhaique, Chile, Patagonia, South America, January.
00451586
Tui De Roy / Black-backed Jackal (Canis mesomelas) trying to scare away Ruppell's Griffons (Gyps rueppellii) from male Grant's Gazelle (Nanger granti) carcass, Ol Pejeta Conservancy, Kenya
00450419
Michael Quinton / Dall's Sheep (Ovis dalli) ram running, Alaska
00451924
Yva Momatiuk & John Eastcott / Sockeye Salmon (Oncorhynchus nerka) male and females during spawning run, Adams River, Roderick Haig-Brown Provincial Park, British Columbia, Canada
00451918
Yva Momatiuk & John Eastcott / Sockeye Salmon (Oncorhynchus nerka) male and female swimming in small stream during spawning run, Roderick Haig-Brown Provincial Park, British Columbia, Canada
01315005
Laurent Geslin / Male Roe deer (Capreolus capreolus) poised to run, in tall grass near a factory in Switzerland.
00447043
Donald M. Jones / Ring-necked Pheasant (Phasianus colchicus) male running on prairie, western Montana
01280756
Patricio Robles Gil / Dead male Pink salmon (Oncorhynchus gorbuscha) on river bed, after the run, South Kamchatka, Russia
01275130
GEORGETTE DOUWMA / Cockerel (Gallus gallus domesticus) crowing in hen run, UK
01275128
GEORGETTE DOUWMA / Cockerel (Gallus gallus domesticus) in hen run, UK
01275127
GEORGETTE DOUWMA / Cockerel (Gallus gallus domesticus) in hen run, UK
00440009
Richard Du Toit / Impala (Aepyceros melampus) male running, Chobe National Park, Botswana
00439488
Michael Quinton / Dall Sheep (Ovis dalli) ram, Yukon Territory, Canada
00437476
Michael Quinton / Grizzly Bear (Ursus arctos horribilis) mother and yearling cubs running away from male standing on the ridge, Alaska
00438355
Suzi Eszterhas / Chimpanzee (Pan troglodytes) male running, western Uganda
00432396
Donald M. Jones / Timber Wolf (Canis lupus) group running down American Elk (Cervus elaphus nelsoni) bull, Yellowstone National Park, Wyoming
00432395
Donald M. Jones / Timber Wolf (Canis lupus) group running down American Elk (Cervus elaphus nelsoni) bull, Yellowstone National Park, Wyoming
