Colin Varndell / Adult Adder (Vipera berus) basking on pebble beach. Abbotsbury, Dorset, UK, March.

Charlie Summers / Group of Rockhopper Penguins (Eudyptes chrysocome) making their way from the nesting colony to the sea. Pebble Island, Falkland Islands

Miles Barton / Panamanian golden frog (Atelopus ziteki) reacting to the image of a frog on a video monitor. Experiment carried out to reveal the purpose of the frog's waving behaviour - to attract mates and deter ri...

Sue Flood / Abandoned Russian research base, Hooker Island, Franz Josef Land, Russian Arctic, July 2007

Poinsignon and Hackel / Common kingfisher {Alcedo atthis} diving underwater to take fish, Lorraine, France

Jack Dykinga / Virgin river flowing through Zion Canyon Narrows, with reflections of afternoon sun, Zion National Park, Utah, USA

Wild Wonders of Europe / Lilja / Petra tou Romiou (Aphrodite's Rock) Pissouri Bay pebble beach, near Paphos, Cyprus, March 2009

Pete Oxford / Gentoo penguin (Pygoscelis papua) jumping out of the sea to land on beach, Pebble Island, Falkland Islands

Niall Benvie / Pebble encased in ice (studio)

Pete Oxford / Rockhopper Penguin (Eudyptes chrysocome) trio, Pebble Island, Falkland Islands

Pete Oxford / Rockhopper Penguin (Eudyptes chrysocome) jumping, Pebble Island, Falkland Islands

Kirkendall-Spring / Waves surging onto a pebble beach, Monterey Coast, California, USA