Anup Shah / Western chimpanzee (Pan troglodytes verus) males 'Jeje' aged 13 years, 'Tua' aged 53 years and 'Peley' aged 12 years grooming, Bossou Forest, Mont Nimba, Guinea. January 2011.

Anup Shah / Olive baboon (Papio cynocephalus anubis) female grooming another females baby, Masai Mara National Reserve, Kenya, August

Anup Shah / Olive baboon (Papio cynocephalus anubis) female with baby aged 1-3 months being groomed, Masai Mara National Reserve, Kenya, September

Florian Möllers / Quinling Golden snub nosed monkey (Rhinopitecus roxellana qinlingensis), family group resting and grooming in a stand of trees. Zhouzhi Nature Reserve, Qinling Mountains, Shaanxi, China.

Florian Möllers / Quinling Golden snub nosed Monkey (Rhinopitecus roxellana qinligensis), two females with young grooming. Zhouzhi Nature Reserve, Qinling Mountains, Shaanxi, China.

Uri Golman / Asiatic lioness (Panthera leo persica) licking another lioness from the pride, Gir Forest NP, Gujarat, India

Anup Shah / Black backed jackals (Canis mesomelas) one pup grooming another, Masai Mara National Reserve, Kenya

Florian Möllers / Golden snub-nosed monkey (Rhinopithecus roxellana qinlingensis) females, subadults and infants socialising in tree, Zhouzi Nature Reserve, Qinling mountains, Shaanxi, China. April 2006

Anup Shah / Olive baboons (Papio cynocephalus anubis) grooming one another, with baby sitting alongside adults. Masai Mara National Reserve, Kenya. March.

Anup Shah / Olive baboon (Papio cynocephalus anubis) grooming a male baby aged 3-6 months Masai Mara National Reserve, Kenya. March.

Chris Gomersall / King penguins (Aptenodytes patagonicus) allo-preening. Fortuna Bay, South Georgia, Antarctica.

Anup Shah / Chimpanzee {Pan troglodytes} grooming companion Sweetwater Sanctuary, Mahesa, Kenya