Luis Quinta / Ghost crab (Ocypode africana) on beach, Island of Principe, UNESCO Biosphere Reserve, Democratic Republic of Sao Tome and Principe, Gulf of Guinea

Staffan Widstrand / Horned ghost crab (Ocypode ceratophthalma) Lowland rainforest, Karawawi River, Kumawa Peninsula, mainland New Guinea, Western Papua, Indonesian New Guinea, on the Science et Images "Expedition Papua,...

Ingo Arndt / Horn-eyed ghost crab (Ocypode ceratophthalma) camouflaged in the sand, emerging from burrow, Christmas Island, Indian Ocean, Australian Territory

Alex Mustard / A trail of sand balls left behind on the beach after the feeding of a ghost crab (Ocypode sp.). The beach is rich in olivine mineral, which is why it is green. Floreana Island, Galapagos Islands, Ecua...

Lynn M. Stone / Atlantic Ghost Crab (Ocypode quadrata) female with egg cluster tucked underneath, making its way to lay eggs, Pinellas County, Florida, USA.

Steve Gettle / Painted Ghost Crab (Ocypode gaudichaudii), Costa Rica

Steve Gettle / Painted Ghost Crab (Ocypode gaudichaudii) emerging from burrow, Costa Rica

Lorraine Bennery / Crab (Ocypode sp.) on beach, Western coastline, Madagascar.

Lorraine Bennery / Crab (Ocypode sp.) on beach, Western coastline, Madagascar.

Lorraine Bennery / Crab (Ocypode sp.) on beach, Western coastline, Madagascar.

Sebastian Kennerknecht / Ghost Crab (Ocypode saratan) on beach, Salalah, Oman

Doug Wechsler / Ghost crab (Ocypode quadrata) eating Lady crab (Ovalipes ocellatus) Sea Isle City, New Jersey, USA, June.