Frederique Olivier / Group of Adelie penguins (Pygoscelis adeliae) walking away from the camera on sea ice at sunset, with one solitary penguin in the foreground, Antarctica.

Ingo Arndt / Grizzly bear (Ursus arctos horribilis) silhouette at sunset, Lake Clark National Park, Alaska, USA, July.

Will Burrard-Lucas / Lion (Panthera leo) walking over the plain, taken with remote camera. Serengeti National Park, Tanzania. December

Mark Hamblin / Raven (Corvus corax) adult calling from rock in mountain habitat, Scotland, UK. November. Third place in British Birds Bird Photograph of the Year competition 2016.

Ann & Steve Toon / Spring wildflowers, Papkuilsfontein farm, Nieuwoudtville, Northern Cape, South Africa, September 2015

Wim van den Heever / Lioness (Panthera leo) sitting during misty sunrise, Okavango Delta, Botswana.

Yva Momatiuk & John Eastcott / Climber climbing rock, Indian Creek, Utah

Matthias Breiter / Bald Eagle (Haliaeetus leucocephalus) on iceberg, LeConte Bay, Tongass National Forest, Alaska

Wim van den Heever / Pied avocet (Recurvirostra avosetta) feeding in shallow water at dawn, Etosha National Park, Namibia.

Brent Stephenson / Young Polar bear (Ursus maritimus) walking across melting sea ice, Scott Inlet, Baffin Island, Canadian Arctic, August. Vulnerable species.

Eric Baccega / African elephant (Loxodonta africana) and black rhinoceros (Diceros bicornis) bathing at waterhole at night with moon, Okaukuejo, Etosha National Park, Namibia, Africa. Taken with infrared camera.

Wim van den Heever / Bottlenosed dolphin (Tursiops truncatus) porpoising during annual sardine run, Port St Johns, South Africa. June.