Ben Hall / Red deer (Cervus elaphus) stags fighting at dusk during rutting season, Cheshire, UK October

Ben Hall / Shelducks (Tadorna tadorna) pair in flight at dusk, Lancashire, UK

Ben Hall / Red grouse (Lagopus lagopus scotica) silhouetted on gritstone edge at dawn, Derbyshire, UK February

Sergey Gorshkov / Giraffe (Giraffa camelopardalis) three standing together, silhouetted at dusk, Okavango Delta, Botswana

Michael W. Richards / Domesticated Indian elephants (Elephas maximus) being walked to forest at sunset, Pench Tiger Reserve, Madhya Pradesh, India

Andrew Parkinson / Coot (Fulica atra) silhouetted as it descends a grass bank, Derbyshire, UK, March. Highly commended in the Portfolio category of the BWPA Competition 2016.

Andrew Parkinson / Red squirrel (Sciurus vulgaris) adult backlit by winter sunlight, Formby Point, Lancashire, UK, December.

Andy Rouse / Silhouetted Grizzly Bear (Ursus arctos horribilis) walking. Katmai, Alaska, USA, September.

Edwin Giesbers / Long-tailed / Crab-eating macaque (Macaca fascicularis) feeding in darkness before dawn, Bako National Park, Sarawak, Borneo, Malaysia

Andy Rouse / African lion (Panthera leo) outline breathing at dawn, Kenya

Richard Du Toit / Chacma baboon backlit at dawn {Papio ursinus} Chobe NP, Botswana

Doug Perrine / Bottlenose dolphin {Tursiops truncatus} Cayman Islands, Caribbean (Non-ex).