Glenn Bartley / Chestnut-mandibled Toucan (Ramphastos swainsonii), Costa Rica

Claudio Contreras / Keel billed toucan (Ramphastos sulfuratus) calling, captive.

Claudio Contreras / Keel billed toucan (Ramphastos sulfuratus), captive.

Bence Mate / Keel billed toucan (Ramphastos sulfuratus) in aggressive encounter with Montezuma Oropendola (Psarocolius montezuma) in rain, Laguna del Lagarto, Santa Rita, Costa Rica. Front cover bookplate of the H...

Nick Garbutt / Toco Toucans (Ramphastos toco) taking flight from the forest canopy. Banks of the Cuiaba River, northern Pantanal, Mato Grosso, Brazil.

Nick Garbutt / Toco Toucan (Ramphastos toco) feeding on fruit in forest canopy. Banks of the Cuiaba River, northern Pantanal, Mato Grosso, Brazil.

Sue Flood / Chestnut mandibled toucan (Ramphastos ambiguus swainsonii) near Boca Tapada, Costa Rica. January 2011

Luiz Claudio Marigo / Channel-billed / Ariel toucan (Ramphastos vittelinus ariel) perched in flowering tree in city street, Rio de Janeiro city, Southeastern Brazil.

Nick Garbutt / A pair of Toco Toucans (Ramphastos toco) flying across the Piquiri River at dawn, northern Pantanal, Mato Grosso, Brazil, September. (Digitally Modified)

Bence Mate / Keel-billed Toucan (Ramphastos sulfuratus) attacking a Brown-hooded Parrot (Pionopsitta haematotis) Laguna del Lagarto, Santa Rita, Costa Rica

Bence Mate / Keel-billed Toucan (Ramphastos sulfuratus) in aggressive encounter with a Montezuma Oropendola (Psarocolius montezuma), Laguna del Lagarto, Santa Rita, Costa Rica

Mary McDonald / Chestnut-mandibled Toucan (Ramphastos ambiguus swainsonii) adult bird perched on the edge of the forest. Laguana del Lagarto, Costa Rica, Central America.