Jim Brandenburg / Arctic Wolf (Canis lupus) alpha male known as Buster, smelling Arctic Poppy (Papaver lapponicum) flowers, Ellesmere Island, Nunavut, Canada
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Jim Brandenburg / Timber Wolf (Canis lupus) alpha pair in light snowfall, Minnesota
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Justine Evans / White handed gibbon, alpha male hanging in tree {Hylobates lar} Khao Yai NP, Thailand 'Cassius'
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Anup Shah / Chimpanzees gather round alpha male 'Frodo' in tree feeding on dead Bushbuck fawn {Pan troglodytes schweinfurtheii} Gombe NP, Tanzania. 2002
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Anup Shah / Chimpanzees gather round alpha male 'Frodo' in tree with dead Bushbuck fawn {Pan troglodytes schweinfurtheii} Gombe NP, Tanzania. 2002
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Anup Shah / Alpha male 'Frodo' displays 'keep away' message to begging chimps wanting share of Bushbuck fawn prey {Pan troglodytes schweinfurtheii}, Kasekela community, Gombe NP, Tanzania. 2002
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Anup Shah / Alpha male Chimpanzee 'Frodo', 26-years-old {Pan troglodytes schweinfurtheii} Kasekela community, Gombe NP, Tanzania. 2002
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Anup Shah / Ousted alpha male Chimpanzee 'Freud' feeding on mangoes, {Pan troglodytes schweinfurtheii} Kasekela community, Gombe NP, Tanzania. 2002
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Anup Shah / Eastern chimpanzee (Pan troglodytes schweinfurtheii) 'Alpha' male 'Ferdinand' aged 21 years walking portrait . Gombe National Park, Tanzania. May 2014.
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Anup Shah / Eastern chimpanzee (Pan troglodytes schweinfurtheii) female 'Schweini' aged 23 years apologising to 'Alpha' male 'Ferdinand' aged 21 years . Gombe National Park, Tanzania. May...
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Anup Shah / Eastern chimpanzee (Pan troglodytes schweinfurtheii) 'Alpha' male 'Ferdinand' aged 21 years walking portrait . Gombe National Park, Tanzania. May 2014.
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Anup Shah / Eastern chimpanzee (Pan troglodytes schweinfurtheii) alpha male 'Ferdinand' aged 22 years feeding on the honeycomb from a bees nest watched by his brother 'Faustino' aged 25 years . Go...
