Bence Mate / Close up of head and prothorax of male Elephant beetle (Megasoma Elephas) with horns, Santa Rita, Costa Rica

Roland Seitre / Silver Jewel Scarab Beetle (Chrysina optima) on red petals. Poas Volcano National Park, Caribbean Coast, Vara Branca, Costa Rica.

John Abbott / Rainbow scarab beetle (Phanaeus difformis) male, Texas, USA

John Abbott / Eastern Green june beetle (Cotinis nitida) in flight, Texas, USA, October

Tony Phelps / Green-grooved Dung Beetle (Scarabaeus rugosus) male with dung ball. deHoop NR, Western Cape, South Africa

Solvin Zankl / Rhinoceros beetle (Oryctes nasicornis) on branch of tree, Germany

Jouan & Rius / Hercules Rhinoceros beetle (Dynastes hercules), tropical forest habitat, Costa Rica

Nick Garbutt / Male Three-horned Rhinoceros beetle {Chalcosoma mollenkampi} on buttress root, lowland rain forest, Danum Valley, Sabah, Borneo.

Peter Blackwell / Scarab beetle on dung ball. Kenya, East Africa

Rolf Nussbaumer / Dung beetle rolling dung ball {Canthon sp} Texas, USA.

Mark Moffett / Golden Scarab Beetle (Plusiotis resplendens) portrait, Monteverde Reserve, Costa Rica

Michael & Patricia Fogden / Scarab Beetle on a Guzmania flower, Damaraland, Namibia