John Shaw / Sally lightfoot crabs (Grapsus grapsus) on the beach at Puerto Egas, Santiago Island, Galapagos, May.

Bernard Castelein / Marsh crab (Neosarmatium meinerti) collectiing mangrove tree leaves and carrying to its hole, Morombe, Madagascar

Kerstin Hinze / Land Crab (Neosarmatium africanum) portrait. St Lucia, Indian Ocean coast, eastern South Africa.

Kerstin Hinze / Land Crab (Neosarmatium africanum) portrait. St Lucia, Indian Ocean coast, eastern South Africa.

Bernard Castelein / Marsh Crab (Neosarmatium meinerti) Zanzibar, Tanzania

Marguerite Smits Van Oyen / Red ghost crabs {Neosarmatium meinerti} pull mangrove leaf into hole to eat. East Africa