Mark Moffett / Weaver Ant (Oecophylla longinoda) group grab an adjacent leaf and stem with mandibles and toes and pull, gradually binding them together, Malaysia

Joao Burini / Army ant (Eciton burchellii) major soldier, Parque da Onca Parda Private Reserve, Sao Paulo, Atlantic Forest South-East Reserves, UNESCO World Heritage Site, Brazil.

Emanuele Biggi / Golden green stag beetle (Lamprima sp.), adult male with big mandibles and a shiny iridescent coloration, Italy. Captive.

Emanuele Biggi / Stag beetle (Prosopocoilus spp.) close up of manibles, occurs in Southeast Asia.

Emanuele Biggi / Weaver ant (Oecophylla) carrying pupa with its mandibles to a new nest. Captive.

Paul Bertner / Long-horned Beetle (Aegosoma ossea) with large mandibles, Danum Valley Conservation Area, Sabah, Borneo, Malaysia

Stephen Dalton / Snail track showing mandibles on algae, Rookery Wood, Sussex, England, UK, May.

Steve Downer / Zebra jumping spider (Salticus scenicus) grooming mandibles with maxillary palps on dandelion seed head, UK. May.

Steve Downer / Zebra jumping spider (Salticus scenicus) grooming mandibles with maxillary palps on dandelion seed head, UK.

Steve Downer / Close up of a Desert locust (Schistocerca gregaria) feeding on grass, controlled conditions.

Alex Hyde / Giant forest ant (Camponotus gigas) soldier. Maliau Basin, Sabah, Borneo.

Alex Hyde / Giant forest ant (Camponotus gigas) Danum Valley, Sabah, Borneo.