Markus Varesvuo / Black Grouse (Tetrao tetrix) two males fighting, Liminka, Finland, March

Markus Varesvuo / Black Grouse (Tetrao tetrix) two males fighting, Liminka, Finland, March

Ammonite / African lioness (Panthera leo) growling and swatting at a sub-adult male approaching a kill, footage taken at night using thermal camera technology, Masai Mara, Kenya.

Roland Seitre / Eastern Wolf (Canis lupus) showing submissive behavior to dominant individual, Omega Park, Montebello, Quebec, Canada

Laurent Geslin / Black backed jackals fighting {Canis mesomelas} Skeleton Coast, Namibia

Laurent Geslin / Black backed jackal, submission {Canis mesomelas} Skeleton Coast, Namibia

Laurent Geslin / Two male urban Red foxes, territorial posture at junction of territories {Vulpes vulpes} London, UK

Laurent Geslin / Two male urban Red foxes, mouths open, one in submissive posture {Vulpes vulpes} London, UK

Laurent Geslin / Two dominant male urban Red foxes fighting on road {Vulpes vulpes} London, UK

Laurent Geslin / Two dominant male urban Red foxes fighting on road, mouths open {Vulpes vulpes} London, UK

Laurent Geslin / Two dominant male urban Red foxes fighting on road {Vulpes vulpes} London, UK

Bence Mate / Grey heron (Ardea cinerea) standing on the neck of another Grey heron, Lake Csaj, Kiskunsagi National Park, Hungary