Andrew Parkinson / Pair of Great skuas (Stercorarius skua) on a misty moorland with person walking close to their territory. Shetland Islands, Scotland, UK, June. (non-ex)

Floris van Breugel / Twilight descends on photographer's mountain camp with light inside tent, in Oregons highest mountain forest on Drake Peak at 8,500 feet. Oregon, USA, January 2010.

Michel Poinsignon / Flock of Mute swans (Cygnus olor) in winter, snowing, Lorraine, France, February

Floris van Breugel / Ponderosa Pine trees (Pinus ponderosa) coated in frost, during heavy snowfall, Deschutes National Forest, Oregon, USA, December 2009

Floris van Breugel / View of Pacific coastline in fog and mist, Ruby Beach, Olympic National Park, Washington, USA, June 2008

Philip Stephen / Huge wave enveloping fishing vessel "Harvester" while operating in the North Sea. Europe, November 2010. Property released.

Christophe Courteau / Clouds during the rainy season in the Kalahari Desert, Botswana, March 2009

Steve Knell / Boreray, under low cloud cover. This holds the largest gannetry in UK (60,000 pairs) St. Kilda, Scotland, July 2009

Philippe Clement / Lightning during thunderstorm above petrochemical industry seen through magnifying glass held against illuminated globe of the earth Digital composite

Philippe Clement / Lightning during thunderstorm above petrochemical industry inside incandescent lamp / bulb against black background. Digital composite

Nick Upton / 2020VISION / Common / Eurasian cranes (Grus grus) small flock of juveniles taking flight over the frozen, snow covered pastureland on a foggy winter morning. These Cranes were released by the Great Crane Project...

Nick Garbutt / Aloe tree at dawn. Anjampolo Forest, southern Madagascar.

Orsolya Haarberg / Musk ox (Ovibos moschatus) snowed in in snow storm, Dovre-Sunndalsfjella National Park, Sor-Trondelag, Norway, March, HIGHLY COMMENDED: Animals in their Environment, Wildlife Photographer of the Year...

Peter Cairns / Flooded birch and alder woodland in autumn, Cairngorms National Park, Scotland, UK, November, RUNNER UP: In Praise of Plants category of the Wildlife Photographer of the Year Competition 2010

David Woodfall / Severn Estuary at ebb tide in rainstorm. This habitat is potentially threatened by proposed tidal barrage. England, UK April 2010

Pete Oxford / Forest vegetation in mist on the western slope of the Mindo Cloud Forest, Ecuador, South America

Stephen Dalton / Owl butterfly (Caligo eurilochus) sheltering from rain

David Tipling / Brown hare (Lepus europaeus) sitting out blizzard in stubble field, Norfolk, England January

David Woodfall / Drought affected agricultural land, monsoon wind blowing top soil from fields, Madhya Pradesh, India, Asia

David Woodfall / Man standing outside building in pouring rain, Ganges delta, Bangladesh, November 2008

Uri Golman / Photographer Uri Golman on cross country skies in snow blizzard, Scoresbysund, North East Greenland, March 2009. Model released

Paul Hobson / Signal Crayfish {Pacifastacus leniusculus} Leicestershire, UK

Philip Dalton / Waterspout over Lake Victoria, Uganda

Philip Dalton / Waterspout over Lake Victoria, Uganda

Inaki Relanzon / Motorcyclist in heavy rain carryaing umbrella, Nyaungshwe, Shan State, Myanmar, Burma August 2009

Sven Zacek / Snowy owl (Bubo scandiaca) perched on tree stump, in snow, Ontario, Canada.

Edwin Giesbers / Common frog (Rana temporaria) sheltering from rain under toadstool (Macrolepiota procera) the Netherlands

Dave Bevan / Long eared owl (Asio otus) perched on post, shaking wings in rain, Wales, UK, captive

Chris Gomersall / Golden eagle (Aquila chrysaetos) sub-adult in winter snow shower, captive bird, Speyside, Scotland, February

Sue Flood / Aerial view of iceberg, with majority of berg visible under the water, Antarctic peninsula, February 2007

George Sanker / Moose (Alces alces) bull walking through woods during a snowstorm, Grand Teton National Park, Wyoming, USA, October

Andrew Walmsley / Male House sparrow (Passer domesticus) peched on a fallen Fir tree in heavy rain, Christchurch, New Zealand, March

Poinsignon and Hackel / Sillhouette of Black woodpecker {Dryocopus martius} flying from nest hole in tree trunk in mist / rain, ancient forest, Vosges mountains, Lorraine, France. Overall winner and winner of the Birds categ...

David Woodfall / Arctic terns (Sterna paradisaea) in flight with a researcher walking, in mist, Northumberland, UK, June 2009

Alex Hyde / Storm clouds over Back Tor, Peak District National Park, England, UK. March 2008.

Andy Rouse / Aerial view of ice ridges at front of Peters Glacier, King Haakon Bay, South Georgia, Antarctica, December 2006

Andy Rouse / Aerial view of Peters Glacier, King Haakon Bay, South Georgia, Antarctica, December 2006

Andy Rouse / Masai giraffe (Giraffa camelopardalis tippelski) showing long tongue, captive, Marwell Zoo, UK

David Tipling / People watching flock of Japanese / Red crowned cranes {Grus japonensis} Akan, Hokkaido, Japan, February 2008

David Tipling / King Penguin {Aptenodytes patagonicus} colony huddled together during storm, Right Whale Bay, South Georgia, November

David Tipling / King Penguin {Aptenodytes patagonicus} colony huddled together during storm, Right Whale Bay, South Georgia, November

David Tipling / King Penguin {Aptenodytes patagonicus} colony huddled together during storm, Right Whale Bay, South Georgia, November

Ross Hoddinott / Widemouth bay beach and coastline at low tide with dark clouds, north Cornwall, UK. May 2009.

Edwin Giesbers / Siberian tiger (Panthera tigris altaica) walking in snowstorm, captive

Andy Rouse / Common wildebeest (Connochaetes taurinus) crossing river on migration in rain, Masai Mara, Kenya, Africa

David Tipling / King Penguin {Aptenodytes patagonicus} colony huddled together during blizzard, Right Whale Bay, South Georgia, November

Roy Mangersnes / Man outside cabin in snowstorm, Agardh Bay, East coast of Spitsbergen, Svalbard, Norway, March 2009. Model released

Mark Bowler / Peruvian red uakari monkey (Cacajao calvus ucayalii) eating aguaje palm fruits (Mauritia flexuosa) Yavari River, Peru, vulnerable species, wild

Jack Dykinga / Pinyon pine (Pinus edulis) and Utah juniper (Juniperus osteosperma) on kaibab limestone spires in morning fog, North Rim, North Kalbab Trail, Grand Canyon National Park, Arizona, USA

Andy Rouse / European Robin (Erithacus rubecula) in snow, Wales, UK

Andy Rouse / Walrus (Odobenus rosmarus) pair on ice in landscape, Svalbard, Norway

Jeff Vanuga / Two male Elk {Cervus canadensis} in velvet, resting in foggy meadow at dawn, Yellowstone National Park, Wyoming, USA, July

Paul Hobson / Hoar frost covering landscape, Peak District, UK, New years day 2009

Paul Hobson / Two trees covered in hoar frost, Peak District, UK, New years day 2009

Mark Taylor / Tricolour Cavalier King Charles Spaniel puppy, lying with chin on floor.

Edwin Giesbers / Gentoo Penguin {Pygoscelis papua} walking through snow storm, Antarctica

Edwin Giesbers / Gentoo Penguin {Pygoscelis papua} walking through snow storm, Antarctica

Ross Hoddinott / Walker in snow storm, near Okehampton, Devon, UK.

Tim Edwards / Sunrise over Loch Bad a Ghaill, near Inverpolly nature reserve. Ross and Cromarty, Scotland.

Eric Baccega / Polar bear resting in blizzard{Ursus maritimus} Churchill, Manitoba, Canada

Staffan Widstrand / Polar bear {Ursus maritimus} walking during snow strom, Churchill, Manitoba, Canada.

Andrew Parkinson / Hikers climb A Chailleach in blizzard, Monadhliath mtns, Highlands, Scotland.

Pete Cairns / Bison {Bison bison} in snow blizzard. Yellowstone National Park, Wyoming, USA

Anup Shah / Patas monkey {Erythrocebus patas} upending kidnapped baby. Laikipia plateau, Kenya.

Asgeir Helgestad / Winter storm with snow drifting. Norefjell mountains, Buskerud, Norway

George Sanker / Coast of Acadia National Park, Maine during Hurricane Sandy. October, 2012.

George Sanker / Acadia National Park, Maine. The Atlantic Ocean. Post-Hurricane Sandy, October.

George Sanker / Acadia National Park, Maine. The Atlantic Ocean. Hurricane Sandy, October.

George Sanker / Storm surge waves breaking on coastal rocks, with person seen through the sea spray. Acadia National Park, Maine. The Atlantic Ocean, Hurricane Sandy, October 2012.

Denis-Huot / African Elephant (Loxodonta africana) kneeling in mud and rain. Aberdares National Park, Kenya.

Anup Shah / Western chimpanzee (Pan troglodytes verus) infant male 'Flanle' aged 3 years spinning in circles, Bossou Forest, Mont Nimba, Guinea. January 2011.

Ben Hall / Great grey owl (Strix nebulosa) perched in pine tree in snowfall, captive

Mark Carwardine / Whale watching tourist boats overcrowd diving Humpback whale, New England, USA

Fergus Gill / 2020VISION / A misty morning over a mixed woodland in autumn, Kinnoull Hill Woodland Park, Perthshire, Scotland, November 2011. Highly commended in the Wild Woods Category of the British Wildlife Photography Awa...

Philip Stephen / The oil rig J.W McLean buffeted by rough waves during a gale in the North Sea, September 2006

Nick Turner / Two cyclists on The Tarka Trail cycle path on a dark cloudy day, Devon, UK, March 2010.

Ben Hall / 2020VISION / View from the summit of Haystacks towards High Crag, Lake District NP, Cumbria, England, UK, November, with storm sky dark clouds

Chadden Hunter / Cameraman Martyn Colbeck filming King penguin (Aptenodytes patagonicus) chick creche / breeding colony, South Georgia. Taken on location for BBC Frozen Planet series, 2008

Sven Zacek / Roe Deer (Capreolus capreolus) leading herd into heavy snow. Estonia, Europe, January.

Sven Zacek / Sea Kale (Crambe maritima) on a beach with the sun just setting. Estonia, Europe, August 2011.

Nick Garbutt / Severe thunder storm and clouds developing over lowland rainforest in the centre of Maliau Basin. Sabah's 'Lost World', Borneo.

Ole Jorgen Liodden / King Penguin (Aptenodytes patagonicus) chick begging its parent for food, with large colony in background. South Georgia Island, Southern Ocean, November.

Peter Cairns / 2020VISION / Photographer, Peter Cairns' floating hide in blizzard, Loch Insh, Cairngorms NP, Highlands, Scotland, UK, March 2011. 2020VISION Exhibition.

Ole Jorgen Liodden / King Penguin (Aptenodytes patagonicus) in heavy snow fall. South Georgia Islands, Southern Ocean, November.

Ole Jorgen Liodden / King Penguin (Aptenodytes patagonicus) adult and chick at their colony. South Georgia Island, Southern Ocean, November.

Ole Jorgen Liodden / Arctic sunset on mountain peaks. Spitsbergen, Svalbard, September.

Ross Hoddinott / Towanroath Engine House in stormy evening light. Wheal Coates, near St Agnes, Cornwall, UK, July 2011.

Philippe Clement / Lightning during thunderstorm at night over church tower and trees. Belgium, May.

Graham Eaton / Icy Road to 'Worlds End', Esclusham Mountain, in overcast winter weather. Wrexham, Wales, December 2010.

David Noton / Stormy seas at Man O' War Bay, Jurassic Coast, Dorset, England, November 2009.

David Noton / Stormy seas at Man O' War Bay, Jurassic Coast, Dorset, England, November 2009.

David Noton / Stormy seas at Lulworth Cove, Jurassic Coast, Dorset, England, November 2009.

David Noton / Stormy seas at Man O' War Bay, Jurassic Coast, Dorset, England, November 2009.