Kevin Schafer / Emperor Penguin (Aptenodytes forsteri) portrait, Atka Bay, Antarctica

Jerry Monkman / Osprey {Pandion haliaetus} female with chick at nest, Martha's Vineyard, Massachusetts, USA

Jerry Monkman / Sun rising over Lower Makonikey, Martha's Vineyard, Cape Cod, Massachusetts, USA

Thomas Marent / Nymphalid Butterfly (Lebadea martha), Kheaun Sri Nakarin National Park, Thailand

Jan Vermeer / Emperor Penguin (Aptenodytes forsteri) chick, Atka Bay, Antarctica

Jan Vermeer / Emperor Penguin (Aptenodytes forsteri) chick, Atka Bay, Antarctica

Jan Vermeer / Weddell Seal (Leptonychotes weddellii) portrait, Atka Bay, Antarctica

Jan Vermeer / Weddell Seal (Leptonychotes weddellii) scratching nose Atka Bay, Antarctica

Jan Vermeer / Weddell Seal (Leptonychotes weddellii) yawning, Atka Bay, Antarctica

Jan Vermeer / Weddell Seal (Leptonychotes weddellii) portrait, Atka Bay, Antarctica

Jan Vermeer / Weddell Seal (Leptonychotes weddellii) portait, Atka Bay, Antarctica

Jan Vermeer / Weddell Seal (Leptonychotes weddellii) lying on ice, Atka Bay, Antarctica

Jan Vermeer / Weddell Seal (Leptonychotes weddellii) yawning, Atka Bay, Antarctica

Martha Holmes / Bull shark (Carcharhinus leucas) caught on line for sport fishing using grouper as bait, South Andros, Bahamas

Sue Flood / Martha Holmes and Doug Allan eating in tent while filming on location in Canadian Arctic for BBC NHU Blue Planet, May 1998

Ben Osborne / Peter Scoones, Martha Holmes and Adelie Penguin preparing to dive, Antarctica - on location for Life in the Freezer, 1992

Martha Holmes / Cameraman Peter Scoones recording Southern elephant seal on location in Stromness Bay, South Georgia, filming for BBC television series "Life in the Freezer", February 1993

Martha Holmes / Rusty parrotfish {Scarus ferrugineus} male at cleaning station, Red Sea

GEORGETTE DOUWMA / Martha Holmes with drowned juvenile Hawksbill turtle (Eretmochelys imbricata) that had been caught in dicarded fishing net. Andaman Sea, Thailand.

Martha Holmes / Detail of palm leaf, back lit.

Martha Holmes / Dynamited coral reef, Philippines

Martha Holmes / Iceberg, detail of erosion, Antarctica.

Martha Holmes / Tourists viewing Grey glacier, Torres del Paine NP, Chile

Martha Holmes / Cuernos del Paine (2668m) in autumn, Torres del Paine NP, Chile

Martha Holmes / Cuernos del Paine (2668m) in autumn, Torres del Paine NP, Chile

Martha Holmes / Cuernos del Paine (2669m) with walkers, Torres del Paine NP, autumn, Chile.

Martha Holmes / Soft coral {Dendronephthya sp} Red Sea

Martha Holmes / Chameleon (Chamaeleonidae family) on hand, Zambia

Martha Holmes / Chris Packham with southern elephant seal, Seal Island Falkland Islands Sea Lion Island on location for the Really Wild Show, 1994

Martha Holmes / Doug Allan, cameraman, on location for BBC 'Life in the Freezer', Tysfjord, Norway, 1997

Martha Holmes / Lichen {Xanthoria parietina} growing on stone wall, Yorkshire, UK

Martha Holmes / Young chimp chewing bark off stick (Pan troglodytes) Chimfunshi Rehabilitation Centre, Zambia

Martha Holmes / Swimming crab in defensive position {Portunus sebae} Grenadines, Caribbean

Martha Holmes / Iceberg, Antarctica

Martha Holmes / Motor yacht Andando with rainbow in stormy sky, Galapagos Islands

Martha Holmes / Boys in dugout canoe in rain with palm leaf used as umbrella, Papua New Guinea

Martha Holmes / Sailing boat (The Golden Fleece) in ice Antarctica summer

Martha Holmes / Mangrove shoots growing in larva field {R. mangle} Isabella, Galapagos

Martha Holmes / Bow of boat Golden Fleece approaching iceberg, Antarctic Peninsula summer

Martha Holmes / Cameraman Doug Allan in kayak filming Humpback whale for BBC television series Blue Planet, Antarctic Peninsula 1999

Martha Holmes / Paul Atkins filming giant petrel and skua scavenging dead fur seal, Bird Island, South Georgia - for BBC television series Life in the Freezer 1992

MARTHA HOLMES / Keeltail needlefish at night (Platybelone argalus) Dominica, Caribbean

Martha Holmes / Fiery-necked nightjar on ground at night (Caprimulgus pectoralis) South Luangwe NP, Zambia

Martha Holmes / Feather duster worm (Sabellastarte magnifica) on coral head, Grenadines, Caribbean

Martha Holmes / Bleached (Acropora) coral, Papua New Guinea

Mats Forsberg / Producer Martha Holmes and cameraman Martin Saunders inside Texas Bar hut, Svalbard, Norway, on location for Polar Bear film special, 1996

Martha Holmes / Flower coral (Eusmilia fastigiata) with polyps extended, Dominica, Caribbean

Martha Holmes / Rough star coral / brain coral (Isophyllastrea rigida) British Virgin Island, Caribbean

Martha Holmes / Close up of patterns in Brain coral (Colpophyllia breviserialis) British Virgin Island, Caribbean

Martha Holmes / Close up of patterns in Brain coral (Colpophyllia natans) Carriacou, Caribbean

Martha Holmes / Club finger coral (Porites porites) with polyps extended, Jamaica, Caribbean

Martha Holmes / David Rootes, assistant on BBC television series Life in the Freezer, amongst brash ice while helping set up remote cameras to film Adelie penguins swimming, 1992

Martha Holmes / Doug Allan filming Killer whales surfacing at sunset for BBC NHU 'The Blue Planet'. Tysfjord, Norway, 1997

Sue Flood / Martha Holmes, Doug Allan and Nat Kalluk (Inuit guide) in Canadian Arctic, May 1998, while filming for BBC NHU Blue Planet.

Martha Holmes / Cameraman Doug Allan with Inuit guide, filming polar bear swimming, Arctic Canada. On location for BBC Polar Bear programme 1996

Sue Flood / Producer Martha Holmes washing her hair in the Canadian Arctic, May 1998, on location for BBC NHU Blue Planet series

Martha Holmes / Adelie penguin swimming in brash ice (Pygoscelis adeliae) Antarctica

Tui De Roy / Emperor Penguin (Aptenodytes forsteri) group on sea ice, No-Name Rookery, Near Elkstrom Ice Shelf, Princess Martha Coast, Weddell Sea, Antarctica

Tui De Roy / Emperor Penguin (Aptenodytes forsteri) large chick on fast ice, midnight sun in austral spring, No-Name Rookery, Princess Martha Coast, Weddell Sea, Antarctica

Tui De Roy / Emperor Penguin (Aptenodytes forsteri) pair, Ekstrom Ice Shelf, Weddell Sea, Antarctica

Tui De Roy / Emperor Penguin (Aptenodytes forsteri) group of four on sea ice under midnight sun, Ekstrom Ice Shelf, Weddell Sea, Antarctica

Tui De Roy / Emperor Penguin (Aptenodytes forsteri) individual on sea ice in midnight twilight, Ekstrom Ice Shelf, Weddell Sea, Antarctica

Tui De Roy / Emperor Penguin (Aptenodytes forsteri) pair sea ice in midnight twilight, Ekstrom Ice Shelf, Weddell Sea, Antarctica

Tui De Roy / Emperor Penguin (Aptenodytes forsteri) commuting across sea ice to nesting colony, Ekstrom Ice Shelf, Weddell Sea, Antarctica

Tui De Roy / Emperor Penguin (Aptenodytes forsteri) group near Ekstrom Ice Shelf, Weddell Sea, Antarctica

Tui De Roy / Emperor Penguin (Aptenodytes forsteri) walking across ice, Ekstrom Ice Shelf, Weddell Sea, Antarctica

Tui De Roy / Emperor Penguin (Aptenodytes forsteri) on sea ice in midnight twilight, Ekstrom Ice Shelf, Weddell Sea, Antarctica

Tui De Roy / Emperor Penguin (Aptenodytes forsteri) portrait on fast ice edge, Princess Martha Coast, Weddell Sea, Antarctica

Martha Holmes / Aerial view of Victoria Land sea and ice, in summer, Antarctica

Martha Holmes / Staghorn coral with healthy growing white tips {Acropora cervicornis} Grenadine, Caribbean

Martha Holmes / Orphan juvenile Chimpanzee {Pan troglodytes} reaching through bars at rehabilitation centre, Chimfunshi, Zambia

Martha Holmes / Orphan juvenile Chimpanzees {Pan troglodytes} drinking through bars at rehabilitation centre, Chimfunshi, Zambia

Martha Holmes / Captive Chimpanzee {Pan troglodytes} looking through bars of cage, Chimfunshi, Zambia

Martha Holmes / Inuits skinning Caribou (Rangifer tarandus) Wager Bay, NW Territories, Canada, sequence

Martha Holmes / Inuit building igloo on sea ice, Admiralty inlet, Canada, Spring, sequence

Martha Holmes / Black tiger snake (Notechis ater) Bass strait, Chappell Island, Australia

Martha Holmes / Inuits skinning Caribou (Rangifer tarandus) Wager Bay, NW Territories, Canada, sequence

Martha Holmes / Marine iguana male in breeding colours (Amblyrhunchus cristatus) Isabela Island, Galapagos

Martha Holmes / Inuits skinning Caribou (Rangifer tarandus) Wager Bay, NW Territories, Canada, sequence

Martha Holmes / Inuits skinning Caribou (Rangifer tarandus) Wager Bay, NW Territories, Canada, sequence

Martha Holmes / Inuits skinning Caribou (Rangifer tarandus) Wager Bay, NW Territories, Canada, sequence

Martha Holmes / Inuits skinning Caribou (Rangifer tarandus) Wager Bay, NW Territories, Canada, sequence

Martha Holmes / Inuit building igloo on sea ice, Admiralty inlet, Canada, Spring, sequence

Martha Holmes / Inuit building igloo on sea ice, Admiralty inlet, Canada, Spring, sequence

Martha Holmes / Marine iguanas mating (Amblyrhynchus cristatus) Espanola Island, Galapagos

Martha Holmes / Inuits skinning Caribou (Rangifer tarandus) Wager Bay, NW Territories, Canada, sequence

Martha Holmes / Inuit building igloo on sea ice, Admiralty inlet, Canada, Spring, sequence

Martha Holmes / Inuits skinning Caribou (Rangifer tarandus) Wager Bay, NW Territories, Canada, sequence

Martha Holmes / Squirrelfish for sale at Tsukiji fish market Tokyo, Japan. 2000

Martha Holmes / Blue fin tuna meat {Thunnus thynnus} for sale in Tsukiji fish market, Tokyo, Japan. 2000

Martha Holmes / Frozen Bluefin tuna {Thunnus thynnus} at Tsukiji fish market, Tokyo, Japan. 2000

Martha Holmes / Octopus for sale at Tsukiji fish market, Tokyo, Japan. 2000

Martha Holmes / Group of Hippos in river {Hippopotamus amphibius} Luangwa river, Zambia.

Martha Holmes / Crocodiles feeding on dead Hippo {Hippopotamus amphibius} Luangwa River, Zambia.

Martha Holmes / Southern elephant seal {Mirounga leonina} sleeping in sand, close-up of face, Falkland Islands.

Martha Holmes / Southern Elephant Seals {Mirounga leonina} resting in wallow, South Georgia.