Yukihiro Fukuda / Japanese macaque / Snow monkey {Macaca fuscata} female suckling two-week-old baby, Jigokudani, Nagano, Japan

Yukihiro Fukuda / Japanese macaque / Snow monkey {Macaca fuscata} three monkeys basking in the winter sunshine, Jigokudani, Nagano, Japan

Yukihiro Fukuda / Japanese macaque / Snow monkey {Macaca fuscata} male grooming female in courtship in autumn woodland, Jigokudani, Nagano, Japan

Yukihiro Fukuda / Japanese macaque / Snow monkey {Macaca fuscata} mother carrying 8-month baby through heavy snow, Jigokudani, Nagano, Japan

Yukihiro Fukuda / Japanese macaque / Snow monkey {Macaca fuscata} young monkey climbs tree to feed on sprouting leaves in spring, Jigokudani, Nagano, Japan

Yukihiro Fukuda / Japanese macaque / Snow monkey {Macaca fuscata} male examing his foot, Jigokudani, Nagano, Japan

Yukihiro Fukuda / Japanese macaque / Snow monkey {Macaca fuscata} three monkeys basking in the winter sunshine, Jigokudani, Nagano, Japan

Yukihiro Fukuda / Japanese macaque / Snow monkey {Macaca fuscata} feeding on wild flowers in spring, Jigokudani, Nagano, Japan

Yukihiro Fukuda / Japanese macaque / Snow monkey {Macaca fuscata} male grooming female in courtship in autumn, Jigokudani, Nagano, Japan

Yukihiro Fukuda / Japanese macaque / Snow monkey {Macaca fuscata} monkeys basking in the winter sunshine, Jigokudani, Nagano, Japan

Yukihiro Fukuda / Japanese macaque / Snow monkey {Macaca fuscata} two-year-old male, Jigokudani, Nagano, Japan

Yukihiro Fukuda / Japanese macaque / Snow monkey {Macaca fuscata} 7-month-old monkey bathing in hot springs, scratching its head, water at 40 degrees, Jigokudani, Nagano, Japan

Yukihiro Fukuda / Japanese macaque / Snow monkey {Macaca fuscata} pine tree with bark eaten by monkeys to help them survive the long winter, Jigokudani, Nagano, Japan

Yukihiro Fukuda / Japanese macaque / Snow monkey {Macaca fuscata} feeding on wild cherry tree blossom in spring, Jigokudani, Nagano, Japan

Yukihiro Fukuda / Japanese macaque / Snow monkey {Macaca fuscata} young monkey amongst autumn foliage, Jigokudani, Nagano, Japan

Yukihiro Fukuda / Japanese macaque / Snow monkey {Macaca fuscata} female with two-day-old male baby, Jigokudani, Nagano, Japan

Yukihiro Fukuda / Japanese macaque / Snow monkey {Macaca fuscata} young monkeys playing in the sunshine in the trees, Jigokudani, Nagano, Japan

Yukihiro Fukuda / Japanese macaque / Snow monkey {Macaca fuscata} rear view of male's bottom and sex organs, Nagano, Japan

Yukihiro Fukuda / Japanese macaque / Snow monkey {Macaca fuscata} pair mating in tree, Jigokudani, Nagano, Japan

Yukihiro Fukuda / Japanese macaque / Snow monkey {Macaca fuscata} group of monkeys basking on the snow in the winter sunshine, Jigokudani, Nagano, Japan

Yukihiro Fukuda / Japanese macaque / Snow monkey {Macaca fuscata} mother and young monkey huddle together for warmth, Jigokudani, Nagano, Japan

Yukihiro Fukuda / Japanese macaque / Snow monkey {Macaca fuscata} monkeys grooming beside river on hot day in summer, Jigokudani, Nagano, Japan

Yukihiro Fukuda / Japanese macaque / Snow monkey {Macaca fuscata} sitting in trees covered in snow, Jigokudani, Nagano, Japan

Yukihiro Fukuda / Japanese macaque / Snow monkey {Macaca fuscata} baby peering over rock edge, Nagano, Japan

Yukihiro Fukuda / Japanese macaque / Snow monkey {Macaca fuscata} male caring for 8-month-old young, Jigokudani, Nagano, Japan

Yukihiro Fukuda / Japanese macaque / Snow monkey {Macaca fuscata} two young monkeys huddle together for warmth, Jigokudani, Nagano, Japan

Yukihiro Fukuda / Japanese macaque / Snow monkey {Macaca fuscata} two-month-old baby feeding on flower in summer, Jigokudani, Nagano, Japan

Yukihiro Fukuda / Japanese macaque / Snow monkey {Macaca fuscata} monkeys searching for food in river, Jigokudani, Nagano, Japan

Yukihiro Fukuda / Japanese macaque / Snow monkey {Macaca fuscata} 8-month-old monkey bathing in hot springs, water at 40 degrees, Jigokudani, Nagano, Japan

Yukihiro Fukuda / Japanese macaque / Snow monkey {Macaca fuscata} footrprints in the snow, Jigokudani, Nagano, Japan

Yukihiro Fukuda / Japanese macaque / Snow monkey {Macaca fuscata} basking in the sun in spring, Jigokudani, Nagano, Japan

Yukihiro Fukuda / Japanese macaque / Snow monkey {Macaca fuscata} female with young in autumn woodland, Jigokudani, Nagano, Japan

Yukihiro Fukuda / Japanese macaque / Snow monkey {Macaca fuscata} adult female bathing in hot springs, playing with the water in her hands, water at 40 degrees, Jigokudani, Nagano, Japan

Yukihiro Fukuda / Japanese macaque / Snow monkey {Macaca fuscata} monkeys grooming in spring, Jigokudani, Nagano, Japan

Yukihiro Fukuda / Japanese macaque / Snow monkey {Macaca fuscata} close up of digits on right hand, Jigokudani, Nagano, Japan

Yukihiro Fukuda / Japanese macaque / Snow monkey {Macaca fuscata} young monkey feeding on wild flowers, Jigokudani, Nagano, Japan

Yukihiro Fukuda / Japanese macaque / Snow monkey {Macaca fuscata} monkeys basking in the sun in spring, Jigokudani, Nagano, Japan

Yukihiro Fukuda / Japanese macaque / Snow monkey {Macaca fuscata} male walking though snow carrying baby on back, Jigokudani, Nagano, Japan

Yukihiro Fukuda / Japanese macaque / Snow monkey {Macaca fuscata} young monkey climbs tree to feed on sprouting leaves in spring, Jigokudani, Nagano, Japan

Yukihiro Fukuda / Japanese macaque / Snow monkey {Macaca fuscata} pair mating in autumn woodland, Jigokudani, Nagano, Japan

Yukihiro Fukuda / Japanese macaque / Snow monkey {Macaca fuscata} group walking along snow trail in heavy snow, Jigokudani, Nagano, Japan

Yukihiro Fukuda / Japanese macaque / Snow monkey {Macaca fuscata} young monkeys playing in the sunshine in the trees, Jigokudani, Nagano, Japan

Yukihiro Fukuda / Japanese macaque / Snow monkey {Macaca fuscata} mother and baby resting beside river on hot day in summer, Jigokudani, Nagano, Japan

Yukihiro Fukuda / Japanese macaque / Snow monkey {Macaca fuscata} two monkeys bathing in hot springs, water at 40 degrees, Jigokudani, Nagano, Japan

Yukihiro Fukuda / Japanese macaque / Snow monkey {Macaca fuscata} feeding on wild flower in spring, Jigokudani, Nagano, Japan

Yukihiro Fukuda / Japanese macaque / Snow monkey {Macaca fuscata} sitting in tree amongst autumn foliage, Jigokudani, Nagano, Japan

Yukihiro Fukuda / Japanese macaque / Snow monkey {Macaca fuscata} monkeys bask in the sunshine and search for food in the woodland amongst melting snow in spring, Jigokudani, Nagano, Japan

Yukihiro Fukuda / Japanese macaque / Snow monkey {Macaca fuscata} 8-month-old monkey bathing in hot springs, water at 40 degrees, Jigokudani, Nagano, Japan

Yukihiro Fukuda / Japanese macaque / Snow monkey {Macaca fuscata} 7-month-old monkey, Jigokudani, Nagano, Japan

Yukihiro Fukuda / Japanese macaque / Snow monkey {Macaca fuscata} 8-month-old monkey bathing in hot springs, sucking finger, water at 40 degrees, Jigokudani, Nagano, Japan

Yukihiro Fukuda / Japanese macaque / Snow monkey {Macaca fuscata} 8-month-old monkey playing with a ball of snow, Jigokudani, Nagano, Japan

Yukihiro Fukuda / Japanese macaque / Snow monkey {Macaca fuscata} young males play fighting, Jigokudani, Nagano, Japan

Yukihiro Fukuda / Japanese macaque / Snow monkey {Macaca fuscata} four young monkeys playing in the sunshine, Jigokudani, Nagano, Japan

Yukihiro Fukuda / Jigokudani valley in autumn, habitat of Japanese macaque / Snow monkey {Macaca fuscata} Nagano, Japan

Yukihiro Fukuda / Japanese macaque / Snow monkey {Macaca fuscata} two 8-month-old monkeys bathing in hot springs, water at 40 degrees, Jigokudani, Nagano, Japan

Yukihiro Fukuda / Japanese macaque / Snow monkey {Macaca fuscata} adult and baby monkey bathing in hot springs, water at 40 degrees, Jigokudani, Nagano, Japan

Yukihiro Fukuda / Japanese macaque / Snow monkey {Macaca fuscata} two monkeys searching for food in river, Jigokudani, Nagano, Japan

Yukihiro Fukuda / Japanese macaque / Snow monkey {Macaca fuscata} 8-month-old monkey, Jigokudani, Nagano, Japan

Yukihiro Fukuda / Japanese macaque / Snow monkey {Macaca fuscata} five-month-old young searching for food amongst fallen leaves in autumn woodland, Jigokudani, Nagano, Japan

Yukihiro Fukuda / Japanese macaque / Snow monkey {Macaca fuscata} 8-month-old monkey bathing in hot springs, water at 40 degrees, Jigokudani, Nagano, Japan

Yukihiro Fukuda / Japanese macaque / Snow monkey {Macaca fuscata} one-month-old babies playing in spring, Jigokudani, Nagano, Japan

Yukihiro Fukuda / Japanese macaque / Snow monkey {Macaca fuscata} 10-month-old monkey climbing trees, Jigokudani, Nagano, Japan

Yukihiro Fukuda / Japanese macaque / Snow monkey {Macaca fuscata} male in autumn woodland displaying to females, Jigokudani, Nagano, Japan

Yukihiro Fukuda / Japanese macaque / Snow monkey {Macaca fuscata} group of monkeys bathing in hot springs, water at 40 degrees, Jigokudani, Nagano, Japan

Yukihiro Fukuda / Japanese macaque / Snow monkey {Macaca fuscata} 7-month-old monkey yawning, Jigokudani, Nagano, Japan

Yukihiro Fukuda / Japanese macaque / Snow monkey {Macaca fuscata} 8-month-old monkeys playing in the snow, Jigokudani, Nagano, Japan

Yukihiro Fukuda / Japanese macaque / Snow monkey {Macaca fuscata} feeding on wild flowers in spring, Jigokudani, Nagano, Japan

Yukihiro Fukuda / Japanese macaque / Snow monkey {Macaca fuscata} female with two-day-old male baby, Jigokudani, Nagano, Japan

Yukihiro Fukuda / Japanese macaque / Snow monkey {Macaca fuscata} 10-month-old monkey hanging upside-down from branch, Jigokudani, Nagano, Japan

Yukihiro Fukuda / Japanese macaque / Snow monkey {Macaca fuscata} monkey in autumn woodland, Jigokudani, Nagano, Japan

Yukihiro Fukuda / Japanese macaque / Snow monkey {Macaca fuscata} male yawning - threatening display, Jigokudani, Nagano, Japan

Yukihiro Fukuda / Japanese macaque / Snow monkey {Macaca fuscata} group of monkeys resting in trees at dawn, Jigokudani, Nagano, Japan

Yukihiro Fukuda / Japanese macaque / Snow monkey {Macaca fuscata} adult monkeys and young drinking from hot springs, water at 40 degrees, Jigokudani, Nagano, Japan

Yukihiro Fukuda / Japanese macaque / Snow monkey {Macaca fuscata} two-year-old monkeys play in the snow, Jigokudani, Nagano, Japan

Yukihiro Fukuda / Japanese macaque / Snow monkey {Macaca fuscata} male resting in the sunshine, Jigokudani, Nagano, Japan

Yukihiro Fukuda / Japanese macaque / Snow monkey {Macaca fuscata} close up of baby suckling from nipple, Jigokudani, Nagano, Japan

Yukihiro Fukuda / Japanese macaque / Snow monkey {Macaca fuscata} male grooming female in courtship in autumn tree, Jigokudani, Nagano, Japan

Yukihiro Fukuda / Japanese macaque / Snow monkey {Macaca fuscata} young monkeys playing in the sunshine in the trees, Jigokudani, Nagano, Japan

Yukihiro Fukuda / Japanese macaque / Snow monkey {Macaca fuscata} male sitting alone waiting for chance to mate with female in autumn, Jigokudani, Nagano, Japan

Yukihiro Fukuda / Japanese macaque / Snow monkey {Macaca fuscata} female monkey with suckling 7-month-old baby bathing in hot springs, water at 40 degrees, Jigokudani, Nagano, Japan

Yukihiro Fukuda / Japanese macaque / Snow monkey {Macaca fuscata} group of monkeys resting in trees at dawn, Jigokudani, Nagano, Japan

Yukihiro Fukuda / Japanese macaque / Snow monkey {Macaca fuscata} female with two-week-old baby, Jigokudani, Nagano, Japan

Yukihiro Fukuda / Japanese macaque / Snow monkey {Macaca fuscata} two-year-old monkeys play in the snow, Jigokudani, Nagano, Japan

Yukihiro Fukuda / Japanese macaque / Snow monkey {Macaca fuscata} female and young huddle together on rock in autumn woodland, Jigokudani, Nagano, Japan

Yukihiro Fukuda / Japanese macaque / Snow monkey {Macaca fuscata} group of monkeys bathing in hot springs, water at 40 degrees, Jigokudani, Nagano, Japan

Yukihiro Fukuda / Japanese macaque / Snow monkey {Macaca fuscata} young monkey feeding on tree bud, Jigokudani, Nagano, Japan

Yukihiro Fukuda / Japanese macaque / Snow monkey {Macaca fuscata} monkeys bathing in hot springs, some grooming, water at 40 degrees, Jigokudani, Nagano, Japan

Yukihiro Fukuda / Japanese macaque / Snow monkey {Macaca fuscata} group of monkeys bathing in hot springs, water at 40 degrees, Jigokudani, Nagano, Japan

Yukihiro Fukuda / Japanese macaque / Snow monkey {Macaca fuscata} young monkeys play with a stick, Jigokudani, Nagano, Japan

Yukihiro Fukuda / Japanese macaque / Snow monkey {Macaca fuscata} female and young huddle together on rock in autumn woodland, Jigokudani, Nagano, Japan

Yukihiro Fukuda / Japanese macaque / Snow monkey {Macaca fuscata} male in snow, Jigokudani, Nagano, Japan

Yukihiro Fukuda / Japanese macaque / Snow monkey {Macaca fuscata} monkeys feeding on wild cherry blossom in spring, Jigokudani, Nagano, Japan

Yukihiro Fukuda / Japanese macaque / Snow monkey {Macaca fuscata} young monkeys play with snow ball, Jigokudani, Nagano, Japan

Yukihiro Fukuda / Japanese macaque / Snow monkey {Macaca fuscata} feeding on Azalea flowers in spring, Jigokudani, Nagano, Japan

Yukihiro Fukuda / Japanese macaque / Snow monkey {Macaca fuscata} feeding on wild flowers, Jigokudani, Nagano, Japan

Yukihiro Fukuda / Japanese macaque / Snow monkey {Macaca fuscata} 8-month-old monkey bathing in hot springs, water at 40 degrees, Jigokudani, Nagano, Japan