Nick Upton / Kerner alpine poppy (Papaver alpinum kerneri) flower growing on karst limestone mountainside at 1600m, Julian Alps, Triglav National Park, Slovenia, July.
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Nick Upton / Alpine calamint / rock thyme (Acinos alpinus) among karst limestone rocks at 1600m in the Julian Alps, Triglav National Park, Slovenia, July.
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Chien Lee / Land Snail (Amphidromus sp), endemic to limestone karst formations, Bukit Sarang Conservation Area, Bintulu, Borneo, Malaysia
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Chien Lee / Land Snail (Amphidromus sp), endemic to limestone karst formations, Bukit Sarang Conservation Area, Bintulu, Borneo, Malaysia
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Tim Laman / Aerial view of the Wayag Islands, an uninhabited group of uplifted limestone (karst) islands, NW of Waigeo Island. West Papua, Indonesia, April 2007
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Tim Laman / A trio of peaks in the Wayag Islands, an uninhabited group of uplifted limestone (karst) islands, NW of Waigeo Island. West Papua, Indonesia, April 2007
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Tim Laman / View of Wayag Islands, an uninhabited group of uplifted limestone (karst) islands, NW of Waigeo Island. West Papua, Indonesia, April 2007
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Tim Laman / Aerial view of the Wayag Islands, an uninhabited group of uplifted limestone (karst) islands, NW of Waigeo Island. West Papua, Indonesia, April 2007
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Tim Laman / Aerial view of a boat making its way through the Wayag Islands, an uninhabited group of uplifted limestone (karst) islands, NW of Waigeo Island. West Papua, Indonesia, April 2007
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Tim Laman / Aerial view of a boat making its way through the Wayag Islands, an uninhabited group of uplifted limestone (karst) islands, NW of Waigeo Island. West Papua, Indonesia, April 2007
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Nick Upton / Karst limestone mountain peak with slopes painted yellow with Small flowered gorse (Ulex parviflora) bushes, Spanish Pyrenees, Revilla, Huesca, Aragon, Spain. July 2009
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Nick Upton / Castro de Gaviotas (Gull's fort) karst limestone rock archway and sea channel leading to narrow La Canalina beach, near Llanes, Asturias, Spain. July 2009
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Nick Upton / Harebell / Bellflower (Campanula rotundifolia) flowering at 1600m on karst limestone boulder at the Cirque de Gavarnie, Haute Pyrenees, France. July 2009
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Nick Upton / Castro de Gaviotas (Gull's fort) karst limestone rock archway and sea channel leading to narrow La Canalina beach, near Llanes, Asturias, Spain. July 2009
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Nick Upton / Waterfalls, karst limestone cliffs and montane pastureland at the Cirque de Gavarnie, Pyrenees National Park, Haute Pyrenees, France. July 2009
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Nick Upton / Waterfalls, karst limestone cliffs, montane pastureland and upper River Gave at the Cirque de Gavarnie, Pyrenees National Park, Haute Pyrenees, France. July 2009
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Nick Upton / Gavarnie Falls cascade down the karst limestone cliffs of the Cirque de Gavarnie, with spray blown upwards by the wind. Pyrenees National Park, Haute Pyrenees, France, July 2009
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Nick Upton / Gavarnie falls cascade down the karst limestone cliffs of the Cirque de Gavarnie, Pyrenees National Park, Haute Pyrenees, France, July 2009
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Nick Upton / Cuevas del mar (Sea caves) beach with arches carved by the sea through karst limestone rocks. Near Llanes, Asturias, Spain. July 2009 Blue Flag Beach.
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Nick Upton / Cuevas del mar (Sea caves) beach with arches carved by the sea through karst limestone rocks. Near Llanes, Asturias, Spain. July 2009 Blue Flag Beach.
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Nick Upton / Chipeta Alta peak and karst limestone escarpment leading to Sayestico peak, Spanish Pyrenees, above Hecho valley, Huesca, Aragon, Spain. July 2009
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Nick Upton / Tourists canyoning down Rio Aragon Subordan through the Boca del Infierno (Hell's Mouth) gorge, carved by river action through karst limestone rock. Hecho valley, Spanish Pyrenees, Huesca, Aragon, Spa...
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Nick Upton / Asturian mountain domestic cattle (Bos taurus) cow with bell on karst limestone clifftop grassland with Atlantic sea in the background, Near Llanes, Asturias, Spain. July 2009
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Nick Upton / Asturian mountain domestic cattle (Bos taurus) grazing on karst limestone clifftop grassland with Atlantic sea in the background, near Llanes, Asturias, Spain. July 2009
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Nick Upton / Asturian mountain domestic cattle (Bos taurus) grazing on karst limestone clifftop grassland with Atlantic sea in the background, near Llanes, Asturias, Spain. July 2009
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Nick Upton / Anisclo Canyon, Bellos river and eroded karst limestone Mondoto peak, Ordesa and Monte Perdido National Park, Huesca, Aragon, Spain. July 2009
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Wild Wonders of Europe / Hermansen / Karst Landscape, Burren National Park, County Clare, Republic of Ireland, June 2009
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Wild Wonders of Europe / Hermansen / Karst landscape near Black Point, Burren National Park, County Clare, Republic of Ireland, June 2009
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Angelo Gandolfi / Karstic cliffs on the "Green Coast", Cantabria, Spain. July 2007.
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Angelo Gandolfi / Secluded cove amongst karstic rocks on the Green Coast, Cantabria, Spain. Picos de Europa mountains in the distance. July 2008.
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Angelo Gandolfi / Turquoise water and karstic rocks on the Green Coast, Cantabria, Spain. Picos de Europa mountains in the distance. July 2008.
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Angelo Gandolfi / Karstic ground with mossy spring that joins the Ebro river. Canyon del Ebro and Rudron, Castilla y Leon, Spain, 2007.
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Jouan & Rius / Crowned lemur (Eulemur / Lemur coronatus) male on limestone karst, Ankarana Special Reserve, North Madagascar
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Jouan & Rius / Male Crowned lemur (Eulemur coronatus) sitting on limestone karst, Ankarana Special Reserve, Madagascar
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Jouan & Rius / Female Crowned lemur (Eulemur coronatus) on limestone karst, Ankarana Special Reserve, Madagascar
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Jouan & Rius / Woman standing on limestone karst ' tsingy' formations, where walking is difficult, Ankarana Special Reserve, Madagascar
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Jouan & Rius / Male Crowned (Eulemur coronatus) lemur on limestone karst, Ankarana Special Reserve, Madagascar
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Jouan & Rius / Male Crowned lemur (Eulemur coronatus) on limestone karst, Ankarana Special Reserve, Madagascar
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Jouan & Rius / Guide helping woman walking on limestone karst ' tsingy' formations Ankarana Special Reserve, Madagascar
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Jouan & Rius / Person walking inside limestone karst ' tsingy' formations with a rope for help, Bemahara NP, Madagascar
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Michael Pitts / Karst limestone formation Central Highlands Papua New Guinea
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Dan Rees / Crowned lemur {Eulemur coronatus} on limestone tsingy (uplifted coral reef) Ankarana
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Pete Oxford / 'Tsingy' landscape: eroded limestone pinnacles, Madagascar Ankarana special reserve
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Nick Garbutt / Dry deciduous forest growing on limestone karst, Manambolo Gorge, Bemaraha NP western Madagascar.
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Nick Garbutt / Limestone karst ' tsingy' formations in Bemaraha NP, western Madagascar.
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Phil Chapman / Male crowned lemur on limestone karst. Madagascar, Ankarana Special Reserve
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Norbert Wu / Limestone islands that were coral reef habitat, Raja Ampat Islands, Indonesia
