Chien Lee / Graceful Madagascar Ground Gecko (Paroedura gracilis), Marojejy National Park, Madagascar

Alex Mustard / Grey seal (Halichoerus grypus) young seal silhouetted against the sun as it dives after breathing, Lundy Island, Devon, UK, Bristol Channel, August

SCOTLAND: The Big Picture / Atlantic puffin (Fratercula arctica) flapping with sand eels in beak, in evening light, Fair Isle, Shetland, Scotland, UK, June.

Sandra Bartocha / Sun rays shining through European beech woodland (Fagus sylvatica). Serrahn, Muritz-National Park, World Natural Heritage site, Germany, Europe. November 2015.

Neil Aldridge / Two Impalas (Aepyceros melampus) ewes grooming each other in evening light on South Africa's Kruger National Park.

Andy Rouse / Mute swan (Cygnus olor) stretching in morning light, UK, February.

Pete Oxford / Whale shark (Rhincodon typus) and scuba divers seen from below, Cenderawasih Bay, West Papua. Indonesia.

Tony Wu / Juvenile Australian sea lion (Neophoca cinerea) silhouetted against blue skies at Carnac Island, Western Australia

Wim van den Heever / Lion (Panthera leo) photographed at night using a side lit spot light, Sabi Sand Game Reserve, South Africa

Solvin Zankl / English oak tree (Quercus robur) in moonlight, Nauroth, Germany, February.

Wim van den Heever / Lion (Panthera leo) male with scars photographed with side-lit spot light at night. Greater Kruger National Park, South Africa, April.

Wim van den Heever / Plains zebra (Equus quagga) at sunset, Savuti Marsh, Botswana.