Kristel Richard / A Kathiawari mare (Equus caballus) standing in Junagadh National Stud, Gujarat, India.
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Edwin Giesbers / Head portrait of Sumatran tiger (Panthera tigris sumatrae) captive
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Edwin Giesbers / Head portrait of Sumatran tiger (Panthera tigris sumatrae) with face half cast in shadow, with aggressive expression, captive
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Edwin Giesbers / Head portrait of Sumatran tiger (Panthera tigris sumatrae) with face half cast in shadow, and mouth open, captive
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Gary K. Smith / Stonechat (Saxicola torquatus rubicola) male perched on barbed wire, Norfolk, UK, January
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Anup Shah / Lilac-breasted roller (Coracias caudata) perched on the back of a Common zebra (Equus quagga). Masai Mara National Reserve, Kenya. December
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Stephen Dalton / Gulf Fritillary Butterfly (Agraulis vanillae) flying near cactus. Everglades, Florida, USA
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Stephen Dalton / Tiger mimic queen butterfly (Lycorea halia) in flight Venezuela, South America
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Dietmar Nill / Great crested grebe (Podicepes cristatus) adult with chick on its back, Germany
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Dietmar Nill / Hebe tiger moth (Arctia festiva) close up of wings, Germany
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Gerrit Vyn / Dead Barn Owl (Tyto alba) on road, Cimarron National Grassland, Kansas, USA, April.
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Edwin Giesbers / Western diamondback rattlesnake {Crotalus atrox} skin detail, captive, from Texas, USA
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Dave Bevan / Long eared owl (Asio otus) perched on post, shaking wings in rain, Wales, UK, captive
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Dave Bevan / Long eared owl (Asio otus) peering round trunk of larch tree, Wales, UK, captive
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Stephen Dalton / Swallowtail butterfly {Papilio machaon} in flight, UK
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Stephen Dalton / Monarch butterfly (Danaus plexippus) female in flight. Did you know? Monarch buttterflies migrate up to 2500 miles from Mexico to Canada.
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Stephen Dalton / Ithomid butterfly {Dircenna sp} in flight, from Venezuelan cloudforest
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Peter Cairns / naturepl.com / Close up of leg rings on Eurasian Crane (Grus grus), Hornborga, Sweden, April
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Alex Hyde / Rosemary beetle (Chrysolina americana) on white background.
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Alex Hyde / Peacock Butterfly {Inachis io} emerging from chrysalis against white background. Sequence 3/6.
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David Tipling / Burrowing owl (Athene cunicularia) resting, Salton Sea, California, USA, April
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Mark Bowler,Mark Bowler / Ball / Royal python (Python regius) coiled, moving away
Andy Rouse / Red Kite (Milvus milvus) in flight, Wales, UK
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Graham Eaton / Rear view of Gadwall (Anas strepera) drying its wings, London Wetland Centre, UK, August
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Jack Dykinga / Sand dunes with cracked clay marking the last flash flood, Creosote bushes (Larrea tridentata) flowering in the background, Mosquito Flats, Death Valley National Park, California, USA
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Andy Rouse / Blue Tit (Parus caeruleus) perched in snow, Wales, UK, January
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John Downer / Crowned crane (Balearica regulorum) portrait, captive
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Anup Shah / Ring-tailed Lemur (Lemur catta) male wrist-marking the tail, lip lared and ears back, during territorial confrontation. Berenty Private Reserve, Madagascar. Oct 2008.
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Anup Shah / Ring-tailed Lemur (Lemur catta) male scent marking. Berenty Private Reserve, Madagascar. Oct 2008.
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Anup Shah / Ring-tailed Lemur (Lemur catta) female carrying baby and scent marking. Berenty Private Reserve, Madagascar. Oct 2008.
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Wild Wonders of Europe / E Haarberg / Chamois (Rupicapra rupicapra) male marking his territory in rutting season, Gran Paradiso National Park, Italy, November 2008
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Albert Lleal / Microchip and needle used to inject sea turtles for marking and tracking data, Barcelona, Spain
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Anup Shah / Banded mongoose {Mungos mungo} group of pups that have been marked for research, Queen Elizabeth NP, Uganda
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Anup Shah / Banded mongoose {Mungos mungo} group of pups that have been marked for research, Queen Elizabeth NP, Uganda
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Dave Watts / Pyrenean chamois {Rupicapra rupicapra pyrenaica} Pyrenees, France
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Anup Shah / Simien jackal / Ethiopian wolf {Canis simensis} marking territory by scraping, Bale Mountains, Bale NP, Ethiopia
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Dave Watts / Eastern Quoll {Dasyurus viverrinus} on log, dark phase, Tasmania, Australia.
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Nick Garbutt / RF- Morpho butterfly displaying on leaf (Morpho achilles). Amazonia, Ecuador. (This image may be licensed either as rights managed or royalty free.)
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Anup Shah / Savanna with Acacia trees, and line of trees marking seasonal water course. Masai Mara, Kenya East-Africa. End of dry season
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David Tipling / Emperor Penguins (Aptenodytes forsteri) chicks Snow Hill Island, Antarctica, November
David Tipling / Emperor Penguins, (Aptenodytes forsteri), chicks huddled together in a creche to keep warm, Weddell Sea, Antarctica
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Jeff Vanuga / Kirk's Dikdik (Madoqua kirki) marking territory by defecating, Samburu National Reserve, Kenya, Africa.
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Colin Varndell / Red Admiral Butterfly (Vanessa atalanta) on Michaelmas daisy flowers. Dorset, UK, October 2012.
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Peter Cairns / 2020VISION / RF- Puffin (Fratercula arctica) in breeding plumage, Hermaness National Nature Reserve. Unst, Shetland Isles, Scotland, UK. June. (This image may be licensed either as rights managed or royalty free.)
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Wild Wonders of Europe / Widstrand / RF- Fritillary butterfly (Melitea sp.) North Velebit National Park, Velebit Nature Park, Rewilding Europe area, Velebit mountains, Croatia, June. (This image may be licensed either as rights managed o...
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Ross Hoddinott / Marbled White butterfly (Melanargia galathea) at rest, Dunsdon Nature Reserve, Devon, UK. July.
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Chris Gomersall / 2020VISION / Grey seal (Halichoerus grypus) scavenging fish around trawlers in Lerwick harbour, Shetland Isles, Scotland, UK, October 2012.
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Anup Shah / Spotted hyaena (Crocuta crocuta) female scent marking grass, Masai Mara National Reserve, Kenya. March
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Nick Garbutt / Male Coyote (Canis latrans) urinating / scent marking. Hayden Valley, Yellowstone National Park, Wyoming, USA, February.
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Danny Green / Tiger (Panthera tigris tigris) female portrait. Endangered. Bandhavgarh, India, November.
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Andrew Parkinson / Bengal Tigers (Panthera tigris) sub-adults, approximately 17-19 months old, playfighting. Endangered. Bandhavgarh National Park, India. Non-ex. Sequence 1 of 2.
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Andrew Parkinson / Bengal Tiger (Panthera tigris) sub-adult, approximately 17-19 months old, lit by morning light, stalking. Endangered. Bandhavgarh National Park, India. Non-ex.
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Andrew Parkinson / Bengal Tiger (Panthera tigris) six year old adult male marks a tree with his scent by rubbing his neck on it. Endangered. Bandhavgarh National Park, India. Non-ex.
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Andrew Parkinson / Bengal Tiger (Panthera tigris) sub-adult, approximately 17-19 months old, amongst forest vegetation. Endangered. Bandhavgarh National Park, India. Non-ex.
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Christophe Courteau / Cheetah (Acinonyx jubatus) male marking territory on tree in the Kalahari Desert, Botswana
Oriol Alamany / Male African lion (Panthera leo) marking his territory at night, Kruger National Park, Transvaal, South Africa, September.
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Donald M. Jones / Black Bear (Ursus americanus) rubbing back on pine, Jasper National Park, Alberta, Canada
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Vladimir Medvedev / Camera trap image of wild male Siberian tiger (Panthera tigris altaica) scent marking the trunk of a cedar (Cedrus) tree, Lazovsky Zapovednik Nature Reserve, Primorsky Krai, Far East Russia, December...
01403016
Loic Poidevin / Indian rhinoceros (Rhinoceros unicornis) juvenile defacating, Kaziranga National Park, Assam, India
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Anup Shah / Lion (Panthera leo) male marking a tree stump by urinating, Masai Mara National Reserve, Kenya, September
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Terry Whittaker / 2020VISION / European Badger (Meles meles) on the North Downs above Folkestone. Kent, UK, June. Camera trap photo.
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Peter Cairns / 2020VISION / RF- Great Spotted Woodpecker (Dendrocopus major) foraging on birch branch. Cairngorms National Park, Scotland. (This image may be licensed either as rights managed or royalty free.)
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Paul Williams / Capybara (Hydrochoerus hydrochaeris) male adult scent marking, Pantanal, Pocone, Brazil
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Paul Williams / Capybara (Hydrochoerus hydrochaeris) male adult scent marking, Pantanal, Pocone, Brazil
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Paul Williams / Capybara (Hydrochoerus hydrochaeris) male adult scent marking, Pantanal, Pocone, Brazil
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Tui De Roy / Cheetah (Acinonyx jubatus) males scent marking tree, El Karama Ranch, Kenya
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Mark Raycroft / White-tailed Deer (Odocoileus virginianus) buck rubbing antlers, North America
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Mark Raycroft / White-tailed Deer (Odocoileus virginianus) buck rubbing antlers, North America
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Philippe Clement / Nieuwe Sluis, oldest extant cast iron lighthouse in the Netherlands, marking the entrance to the Westerschelde at Breskens, Zeeland, February
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Carol Walker / RF- Wild Mustang horses. McCullough Peaks Herd Area, Northern Wyoming, USA. June (This image may be licensed either as rights managed or royalty free.)
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Michel Poinsignon / Starlings (Sturnus vulgaris), adults perched on branch in winter feeding on apple. Lorraine France. 3rd Prize in the Melvita Nature Images Awards competition 2013.
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Sharon Heald / Black rhinoceros (Diceros bicornis) male spray marking, Etosha National Park, Namibia
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Eric Dragesco / Snow Leopard (Panthera unicia) urine sprayed on rocks as territorial scent marking, Altai Mountains, Mongolia
Edwin Giesbers / RF- Rothschild's giraffe (Giraffa camelopardalis rothschildi) two curious giraffes looking down, captive. (This image may be licensed either as rights managed or royalty free.)
01374301
Edwin Giesbers / RF- Rothschild's giraffe (Giraffa camelopardalis rothschildi) head and neck portrait, captive. (This image may be licensed either as rights managed or royalty free.)
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Alex Mustard / RF- Chain moray eel (Echidna catenata) portrait with mouth open. East End, Grand Cayman, Cayman Islands, British West Indies. Caribbean Sea. (This image may be licensed either as rights managed or roy...
01374163
Steven David Miller / Black-necked stork / Jabiru (Ephippiorhynchus asiaticus) pair displaying on nest with fledgling, male on the left, captive, The Wildlife Habitat, Port Douglas, Queensland, Australia, August
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Hermann Brehm / White rhinoceros (Ceratotherium simum) male marking terretory, Solio GR, Kenya
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Donald M. Jones / Elk (Cervus elaphus) bull with grasses on antlers, National Bison Range, Moise, Montana
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Donald M. Jones / Caribou (Rangifer tarandus) bull rubbing tree on fall colored tundra, Denali National Park, Alaska
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Petra Wegner / Longhaired blue-merle Chihuahua puppy, 17 weeks.
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Sergey Gorshkov / Leopard (Panthera pardus) partial head view of adult resting in tree, Okavango Delta, Botswana
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Mark Raycroft / White-tailed Deer (Odocoileus virginianus) buck branch licking to mark territory, North America
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Mark Raycroft / White-tailed Deer (Odocoileus virginianus) buck rubbing antlers against tree trunk to mark territory, North America
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Mark Raycroft / White-tailed Deer (Odocoileus virginianus) buck branch licking to mark territory, North America
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Mark Raycroft / White-tailed Deer (Odocoileus virginianus) buck rubbing tree trunk to mark territory, North America
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Mark Raycroft / White-tailed Deer (Odocoileus virginianus) buck licking plant to mark territory, North America
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Ann & Steve Toon / Nyala (Tragelaphus angasii) territorial bull scent-marking, Mkhuze game reserve, South Africa, May.
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Edwin Giesbers / RF- Toco toucan (Ramphastos toco) head close-up, captive. (This image may be licensed either as rights managed or royalty free.)
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Edwin Giesbers / RF- Male Indian peafowl (Pavo cristatus) head portrait, captive. (This image may be licensed either as rights managed or royalty free.)
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Edwin Giesbers / RF- Male Indian peafowl (Pavo cristatus) displaying, captive. (This image may be licensed either as rights managed or royalty free.)
