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RF - King penguin (Aptenodytes patagonicus) colony with adults and chicks. Salisbury Plain, South Georgia Island. November. (This image may be licensed either as rights managed or royalty free.)
01634681
Nick Garbutt / RF - King penguin (Aptenodytes patagonicus) colony with adults and chicks. Salisbury Plain, South Georgia Island. November. (This image may be licensed either as rights managed or royalty free.)
RF - Black-browed albatross (Thalassarche melanophris) in flight, snow covered coastline in background. Cooper Bay, South Georgia. November 2018. (This image may be licensed either as rights managed o...
01634680
Nick Garbutt / RF - Black-browed albatross (Thalassarche melanophris) in flight, snow covered coastline in background. Cooper Bay, South Georgia. November 2018. (This image may be licensed either as rights managed o...
RF - Southern elephant seal (Mirounga leonina) pup, portrait. Gold Harbour, South Georgia. November. (This image may be licensed either as rights managed or royalty free.)
01634679
Nick Garbutt / RF - Southern elephant seal (Mirounga leonina) pup, portrait. Gold Harbour, South Georgia. November. (This image may be licensed either as rights managed or royalty free.)
RF - King penguins (Aptenodytes patagonicus), two standing on snow, one sleeping or preening. Grytviken, South Georgia. March. (This image may be licensed either as rights managed or royalty free.)
01634678
Nick Garbutt / RF - King penguins (Aptenodytes patagonicus), two standing on snow, one sleeping or preening. Grytviken, South Georgia. March. (This image may be licensed either as rights managed or royalty free.)
RF - Southern elephant seal (Mirounga leonina), two males fighting in colony amongst King penguins (Aptenodytes patagonicus). St Andrews Bay, South Georgia. November. (This image may be licensed eithe...
01634677
Nick Garbutt / RF - Southern elephant seal (Mirounga leonina), two males fighting in colony amongst King penguins (Aptenodytes patagonicus). St Andrews Bay, South Georgia. November. (This image may be licensed eithe...
RF - Chestnut-crowned laughing thrush (Trochalopteron erythrocephalum) foraging, Singalila National Park, India. (This image may be licensed either as rights managed or royalty free.)
01634676
Nick Garbutt / RF - Chestnut-crowned laughing thrush (Trochalopteron erythrocephalum) foraging, Singalila National Park, India. (This image may be licensed either as rights managed or royalty free.)
RF - King penguin (Aptenodytes patagonicus), four returning to sea. St Andrews Bay, South Georgia. November. (This image may be licensed either as rights managed or royalty free.)
01634675
Nick Garbutt / RF - King penguin (Aptenodytes patagonicus), four returning to sea. St Andrews Bay, South Georgia. November. (This image may be licensed either as rights managed or royalty free.)
RF - Red fox (Vulpes vulpes) foraging in falling snow in winter. Hayden Valley, Yellowstone National Park, USA. February. (This image may be licensed either as rights managed or royalty free.)
01634674
Nick Garbutt / RF - Red fox (Vulpes vulpes) foraging in falling snow in winter. Hayden Valley, Yellowstone National Park, USA. February. (This image may be licensed either as rights managed or royalty free.)
RF - Black-browed albatross (Thalassarche melanophris), two flying in opposite directions, another in background. South Atlantic Ocean between The Falklands Islands and South Georgia. November. (This...
01634673
Nick Garbutt / RF - Black-browed albatross (Thalassarche melanophris), two flying in opposite directions, another in background. South Atlantic Ocean between The Falklands Islands and South Georgia. November. (This...
RF - Coyote (Canis latrans) foraging on ice. Madison Valley, Yellowstone National Park, USA. February. (This image may be licensed either as rights managed or royalty free.)
01634672
Nick Garbutt / RF - Coyote (Canis latrans) foraging on ice. Madison Valley, Yellowstone National Park, USA. February. (This image may be licensed either as rights managed or royalty free.)
RF - Coyote (Canis latrans) sitting on ice, looking down. Madison Valley, Yellowstone National Park, USA. (This image may be licensed either as rights managed or royalty free.)
01634671
Nick Garbutt / RF - Coyote (Canis latrans) sitting on ice, looking down. Madison Valley, Yellowstone National Park, USA. (This image may be licensed either as rights managed or royalty free.)
RF - Western red panda (Ailurus fulgens fulgens) sitting in tree, looking down. Singalila National Park, India / Nepal border. March. (This image may be licensed either as rights managed or royalty fr...
01634670
Nick Garbutt / RF - Western red panda (Ailurus fulgens fulgens) sitting in tree, looking down. Singalila National Park, India / Nepal border. March. (This image may be licensed either as rights managed or royalty fr...
RF - Puma (Puma concolor puma) female walking, near Torres del Paine National Park, Patagonia, Chile. December. (This image may be licensed either as rights managed or royalty free.)
01634669
Nick Garbutt / RF - Puma (Puma concolor puma) female walking, near Torres del Paine National Park, Patagonia, Chile. December. (This image may be licensed either as rights managed or royalty free.)
RF - Puma (Puma concolor puma), young male walking in grassland, near Torres del Paine National Park, Patagonia, Chile. December. (This image may be licensed either as rights managed or royalty free.)
01634668
Nick Garbutt / RF - Puma (Puma concolor puma), young male walking in grassland, near Torres del Paine National Park, Patagonia, Chile. December. (This image may be licensed either as rights managed or royalty free.)
RF - Western red panda (Ailurus fulgens fulgens) resting on branch, Singalila National Park, India / Nepal border. (This image may be licensed either as rights managed or royalty free.)
01634667
Nick Garbutt / RF - Western red panda (Ailurus fulgens fulgens) resting on branch, Singalila National Park, India / Nepal border. (This image may be licensed either as rights managed or royalty free.)
Southern elephant seal (Mirounga leonina), two males, equally matched with mouths open in aggression. Out of focus King penguin (Aptenodytes patagonicus) colony in background. Gold Harbour, South Geor...
01634666
Nick Garbutt / Southern elephant seal (Mirounga leonina), two males, equally matched with mouths open in aggression. Out of focus King penguin (Aptenodytes patagonicus) colony in background. Gold Harbour, South Geor...
Towers and Central Massif reflected in Lago Pehoe at sunrise. Torres del Paine National Park, Patagonia, Chile. November 2018.
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Nick Garbutt / Towers and Central Massif reflected in Lago Pehoe at sunrise. Torres del Paine National Park, Patagonia, Chile. November 2018.
Red giant flying squirrel (Petaurista petaurista) gliding between trees at dusk. Sepilok, Sabah, Borneo, Malaysia. May.
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Nick Garbutt / Red giant flying squirrel (Petaurista petaurista) gliding between trees at dusk. Sepilok, Sabah, Borneo, Malaysia. May.
Rafflesia (Rafflesia keithii) flower aged approximately 3 days on rainforest floor. Lower slopes of Mt Kinabalu, Sabah, Borneo, Malaysia.
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Nick Garbutt / Rafflesia (Rafflesia keithii) flower aged approximately 3 days on rainforest floor. Lower slopes of Mt Kinabalu, Sabah, Borneo, Malaysia.
Rafflesia (Rafflesia keithii) flower aged approximately 3 days, on rainforest floor. Lower slopes of Mt Kinabalu, Sabah, Borneo, Malaysia.
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Nick Garbutt / Rafflesia (Rafflesia keithii) flower aged approximately 3 days, on rainforest floor. Lower slopes of Mt Kinabalu, Sabah, Borneo, Malaysia.
King penguin (Aptenodytes patagonicus) displaying, looking skywards, surrounded by immature chicks. Salisbury Plain, South Georgia. November.
01634661
Nick Garbutt / King penguin (Aptenodytes patagonicus) displaying, looking skywards, surrounded by immature chicks. Salisbury Plain, South Georgia. November.
Towers and Central Massif reflected in Laguna Amarga, Torres del Paine National Park, Patagonia, Chile. November 2018.
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Nick Garbutt / Towers and Central Massif reflected in Laguna Amarga, Torres del Paine National Park, Patagonia, Chile. November 2018.
Swallowtail butterfly (Papilio machaon) in flight. North Tyrol, Austria. June.
01634659
Nick Garbutt / Swallowtail butterfly (Papilio machaon) in flight. North Tyrol, Austria. June.
Swallowtail butterfly (Papilio machaon) resting on Thistle (Cirsium sp). North Tyrol, Austria. June.
01634658
Nick Garbutt / Swallowtail butterfly (Papilio machaon) resting on Thistle (Cirsium sp). North Tyrol, Austria. June.
Swallowtail butterfly (Papilio machaon) nectaring on Meadow clary (Salvia pratensis). North Tyrol, Austria. June.
01634657
Nick Garbutt / Swallowtail butterfly (Papilio machaon) nectaring on Meadow clary (Salvia pratensis). North Tyrol, Austria. June.
Alpine rose (Rhododendron ferrugineum) near stream in mountain valley. Fiss, Landeck, North Tyrol, Austria. June 2017.
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Nick Garbutt / Alpine rose (Rhododendron ferrugineum) near stream in mountain valley. Fiss, Landeck, North Tyrol, Austria. June 2017.
Alpine Wildflower meadow, a diversity of species including Meadow clary (Salvia pratensis), Sainfoin (Onobrychis sp) and Rattle (Rhinanthus sp). North Tyrol, Austria. June.
01634655
Nick Garbutt / Alpine Wildflower meadow, a diversity of species including Meadow clary (Salvia pratensis), Sainfoin (Onobrychis sp) and Rattle (Rhinanthus sp). North Tyrol, Austria. June.
Oxeye daisy (Leucanthemum vulgaris) flowers. North Tyrol, Austria. June.
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Nick Garbutt / Oxeye daisy (Leucanthemum vulgaris) flowers. North Tyrol, Austria. June.
Walkway through lowland Dipterocarp (Dipterocarpaceae) rainforest, aerial shot. Danum Valley, Sabah, Borneo, Malaysia. 2018.
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Nick Garbutt / Walkway through lowland Dipterocarp (Dipterocarpaceae) rainforest, aerial shot. Danum Valley, Sabah, Borneo, Malaysia. 2018.
RF - Apollo butterfly (Parnassius apollo) nectaring on flower. North Tyrol, Austria. June. Focus stacked image. (This image may be licensed either as rights managed or royalty free.)
01634691
Nick Garbutt / RF - Apollo butterfly (Parnassius apollo) nectaring on flower. North Tyrol, Austria. June. Focus stacked image. (This image may be licensed either as rights managed or royalty free.)
RF - Rays of sunshine through dead tree trunks along winding river, in snow on misty morning. Firehole Valley. Yellowstone National Park, USA. January (This image may be licensed either as rights mana...
01634689
Nick Garbutt / RF - Rays of sunshine through dead tree trunks along winding river, in snow on misty morning. Firehole Valley. Yellowstone National Park, USA. January (This image may be licensed either as rights mana...
RF - Puma (Puma concolor puma), young male walking on hillside near Torres del Paine National Park, Patagonia, Chile. November. (This image may be licensed either as rights managed or royalty free.)
01634688
Nick Garbutt / RF - Puma (Puma concolor puma), young male walking on hillside near Torres del Paine National Park, Patagonia, Chile. November. (This image may be licensed either as rights managed or royalty free.)
RF - Wildflowers flowering in ancient alpine meadow. North Tyrol, Austria. June. (This image may be licensed either as rights managed or royalty free.)
01634687
Nick Garbutt / RF - Wildflowers flowering in ancient alpine meadow. North Tyrol, Austria. June. (This image may be licensed either as rights managed or royalty free.)
RF - American bison (Bison bison) male standing near Punch Bowl thermal pool, green algae in foreground. Yellowstone National Park, USA. (This image may be licensed either as rights managed or royalty...
01634686
Nick Garbutt / RF - American bison (Bison bison) male standing near Punch Bowl thermal pool, green algae in foreground. Yellowstone National Park, USA. (This image may be licensed either as rights managed or royalty...
RF - Guanaco (Lama guanicoe) walking down slope with mountain towers of Paine in background, Torres del Paine National Park, Patagonia, Chile. December 2018. (This image may be licensed either as righ...
01634685
Nick Garbutt / RF - Guanaco (Lama guanicoe) walking down slope with mountain towers of Paine in background, Torres del Paine National Park, Patagonia, Chile. December 2018. (This image may be licensed either as righ...
RF - Black veined white butterfly (Aporia crataegi) nectaring on Fragrant orchid (Gymnadenia conopsea). North Tyrol, Austria, June. (This image may be licensed either as rights managed or royalty free...
01634684
Nick Garbutt / RF - Black veined white butterfly (Aporia crataegi) nectaring on Fragrant orchid (Gymnadenia conopsea). North Tyrol, Austria, June. (This image may be licensed either as rights managed or royalty free...
RF - Guanaco (Lama guanicoe) standing in front of mountain towers of Paine. Torres del Paine National Park, Patagonia, Chile. December 2018. (This image may be licensed either as rights managed or roy...
01634682
Nick Garbutt / RF - Guanaco (Lama guanicoe) standing in front of mountain towers of Paine. Torres del Paine National Park, Patagonia, Chile. December 2018. (This image may be licensed either as rights managed or roy...
Cinnabar moth (Tyria jacobaeae) caterpillar feeding on Ragwort (Jacobaea vulgaris) in meadow. Wiltshire, England, UK. July.
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Nick Upton / Cinnabar moth (Tyria jacobaeae) caterpillar feeding on Ragwort (Jacobaea vulgaris) in meadow. Wiltshire, England, UK. July.
Goldenrod crab spider (Misumena vatia) female on Field scabious (Knautia arvensis) waiting for prey to land, legs raised, Wiltshire, England, UK. July.
01632502
Nick Upton / Goldenrod crab spider (Misumena vatia) female on Field scabious (Knautia arvensis) waiting for prey to land, legs raised, Wiltshire, England, UK. July.
Eurasian beaver (Castor fiber) female swimming in river. River Otter tributary, Devon, England, UK. June.
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Nick Upton / Eurasian beaver (Castor fiber) female swimming in river. River Otter tributary, Devon, England, UK. June.
European badger (Meles meles) running across garden lawn at night, close to a house. Wiltshire, England, UK. May. Camera trap image.
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Nick Upton / European badger (Meles meles) running across garden lawn at night, close to a house. Wiltshire, England, UK. May. Camera trap image.
Grey squirrel (Sciurus carolinensis) feeding, sitting on garden lawn. Wiltshire, England, UK. May 2019. Camera trap image.
01632392
Nick Upton / Grey squirrel (Sciurus carolinensis) feeding, sitting on garden lawn. Wiltshire, England, UK. May 2019. Camera trap image.
Common frog (Rana temporaria) frogspawn in pond created by damming of woodland stream by Eurasian beaver (Castor fiber). Beaver gnawed branch in background. Devon Beaver Project, Devon Wildlife Trust,...
01632391
Nick Upton / Common frog (Rana temporaria) frogspawn in pond created by damming of woodland stream by Eurasian beaver (Castor fiber). Beaver gnawed branch in background. Devon Beaver Project, Devon Wildlife Trust,...
Common frog (Rana temporaria) frogspawn in pond created by damming of woodland stream by Eurasian beaver (Castor fiber). Beaver gnawed branch in background. Devon Beaver Project, Devon Wildlife Trust,...
01632390
Nick Upton / Common frog (Rana temporaria) frogspawn in pond created by damming of woodland stream by Eurasian beaver (Castor fiber). Beaver gnawed branch in background. Devon Beaver Project, Devon Wildlife Trust,...
Male Common darter dragonfly (Sympetrum striolatum) sun basking on a fence post in early winter sunshine, Rutland Water, Rutland, UK, November.
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Nick Upton / Male Common darter dragonfly (Sympetrum striolatum) sun basking on a fence post in early winter sunshine, Rutland Water, Rutland, UK, November.
Male Common European Crane fly / Daddy long legs (Tipula paludosa) recently emerged and resting on Goose grass / Cleavers (Galium aparine) by a pond, Suffolk, UK, May.
01630541
Nick Upton / Male Common European Crane fly / Daddy long legs (Tipula paludosa) recently emerged and resting on Goose grass / Cleavers (Galium aparine) by a pond, Suffolk, UK, May.
Caddis fly (Trichoptera) resting on riverbank vegetation, Wiltshire, UK, May.
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Nick Upton / Caddis fly (Trichoptera) resting on riverbank vegetation, Wiltshire, UK, May.
Silken cocoon of Narrow-bordered Five-spot Burnet moth (Zygaena lonicerae) attached to grass stem, chalk grassland meadow, Wiltshire, UK, May.
01630539
Nick Upton / Silken cocoon of Narrow-bordered Five-spot Burnet moth (Zygaena lonicerae) attached to grass stem, chalk grassland meadow, Wiltshire, UK, May.
Brown mayfly (Ephemera vulgata) newly emerged, resting on riverside vegetation, River Avon, Lacock, Wiltshire, UK, May.
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Nick Upton / Brown mayfly (Ephemera vulgata) newly emerged, resting on riverside vegetation, River Avon, Lacock, Wiltshire, UK, May.
Brown mayfly (Ephemera vulgata) newly emerged on a riverside nettle leaf, River Avon, Lacock, Wiltshire, UK, May.
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Nick Upton / Brown mayfly (Ephemera vulgata) newly emerged on a riverside nettle leaf, River Avon, Lacock, Wiltshire, UK, May.
Bloody-nosed beetle (Timarcha tenebricosa) in a stand of its food plant Goose grass / Cleavers (Galium aparine) in a chalk grassland meadow, Wiltshire, UK, May.
01630536
Nick Upton / Bloody-nosed beetle (Timarcha tenebricosa) in a stand of its food plant Goose grass / Cleavers (Galium aparine) in a chalk grassland meadow, Wiltshire, UK, May.
Male Black Snipe fly (Chrysopilus cristatus), standing on a Philadelphus leaf in a garden, Wiltshire, UK, May.
01630535
Nick Upton / Male Black Snipe fly (Chrysopilus cristatus), standing on a Philadelphus leaf in a garden, Wiltshire, UK, May.
Black oil beetle (Meloe proscarabaeus) walking on a coast path, seeking a site to dig a burrow for its eggs, north Cornwall, UK, April.
01630534
Nick Upton / Black oil beetle (Meloe proscarabaeus) walking on a coast path, seeking a site to dig a burrow for its eggs, north Cornwall, UK, April.
Black oil beetle (Meloe proscarabaeus) starting to excavate a nest burrow in a footpath on a chalk grassland hillside, the Ridgeway, Wiltshire, UK, April.
01630533
Nick Upton / Black oil beetle (Meloe proscarabaeus) starting to excavate a nest burrow in a footpath on a chalk grassland hillside, the Ridgeway, Wiltshire, UK, April.
Black oil beetle (Meloe proscarabaeus) female excavating a nest burrow in coastal grassland, north Cornwall, UK, April.
01630532
Nick Upton / Black oil beetle (Meloe proscarabaeus) female excavating a nest burrow in coastal grassland, north Cornwall, UK, April.
Beadlet anemone (Actinia equina) in a rock pool, Cornwall, UK, March.
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Nick Upton / Beadlet anemone (Actinia equina) in a rock pool, Cornwall, UK, March.
Alder fly (Sialis sp.) resting on riverside umbel stem, Wiltshire, UK, May.
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Nick Upton / Alder fly (Sialis sp.) resting on riverside umbel stem, Wiltshire, UK, May.
Velvet swimming crab (Necora puber) in a rock pool, hiding among Coralweed (Corallina officinalis), Cornwall, UK, March.
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Nick Upton / Velvet swimming crab (Necora puber) in a rock pool, hiding among Coralweed (Corallina officinalis), Cornwall, UK, March.
Strawberry anemone (Actinia fragracea) in a rock pool alongside Coralweed (Corallina officinalis), Cornwall, UK, March.
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Nick Upton / Strawberry anemone (Actinia fragracea) in a rock pool alongside Coralweed (Corallina officinalis), Cornwall, UK, March.
Spotted longhorn beetle (Rutpela maculata = Strangalia maculata) takes off after feeding on a Common spotted orchid flower (Dactylorhiza fuchsii) in a woodland ride, Lower Woods, Gloucestershire, UK,...
01630527
Nick Upton / Spotted longhorn beetle (Rutpela maculata = Strangalia maculata) takes off after feeding on a Common spotted orchid flower (Dactylorhiza fuchsii) in a woodland ride, Lower Woods, Gloucestershire, UK,...
Common shore crab (Carcinus maenas) hiding among Coralweed (Corallina officinalis) in a rock pool, Cornwall, UK, March.
01630526
Nick Upton / Common shore crab (Carcinus maenas) hiding among Coralweed (Corallina officinalis) in a rock pool, Cornwall, UK, March.
Young female Nursery web spider (Pisaura mirabilis) sun basking on a leaf on a late winter day, Wiltshire garden, UK, February.
01630525
Nick Upton / Young female Nursery web spider (Pisaura mirabilis) sun basking on a leaf on a late winter day, Wiltshire garden, UK, February.
Dense bed of Common mussels (Mytilus edulis) exposed at low tide on rocks fringing a sandy beach, Godrevy, Cornwall, UK, April.
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Nick Upton / Dense bed of Common mussels (Mytilus edulis) exposed at low tide on rocks fringing a sandy beach, Godrevy, Cornwall, UK, April.
Meadow plant bug (Leptopterna dolabrata) female laying an egg on a grass blade in a meadow, Wiltshire, UK, June.
01630523
Nick Upton / Meadow plant bug (Leptopterna dolabrata) female laying an egg on a grass blade in a meadow, Wiltshire, UK, June.
Meadow brown butterfly (Maniola jurtina) nectaring on a Greater knapweed flower (Centaurea scabiosa) in a chalk grassland meadow, Wiltshire, UK, June.
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Nick Upton / Meadow brown butterfly (Maniola jurtina) nectaring on a Greater knapweed flower (Centaurea scabiosa) in a chalk grassland meadow, Wiltshire, UK, June.
Marbled white butterfly (Melanargia galathea) nectaring on a Greater knapweed flower (Centaurea scabiosa) in a chalk grassland meadow, Wiltshire, UK, June.
01630521
Nick Upton / Marbled white butterfly (Melanargia galathea) nectaring on a Greater knapweed flower (Centaurea scabiosa) in a chalk grassland meadow, Wiltshire, UK, June.
Female Kestrel (Falco tinnunculus) soliciting male to mate with her by crouching down on a cliff ledge near their nest site, Cornwall, UK, April.
01630409
Nick Upton / Female Kestrel (Falco tinnunculus) soliciting male to mate with her by crouching down on a cliff ledge near their nest site, Cornwall, UK, April.
Kestrel (Falco tinnunculus) female perched on a cliff ledge while eating a Common shrew (Sorex araneus) brought by her mate as a courtship gift and held in her talons, Cornwall, UK, April.
01630408
Nick Upton / Kestrel (Falco tinnunculus) female perched on a cliff ledge while eating a Common shrew (Sorex araneus) brought by her mate as a courtship gift and held in her talons, Cornwall, UK, April.
Greylag goose (Anser anser) pair swimming with four goslings on a lake, Gloucestershire, UK, April.
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Nick Upton / Greylag goose (Anser anser) pair swimming with four goslings on a lake, Gloucestershire, UK, April.
Kestrel (Falco tinnunculus) male hovering overhead, Cornwall, UK, April.
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Nick Upton / Kestrel (Falco tinnunculus) male hovering overhead, Cornwall, UK, April.
Large red damselfly (Pyrrhosoma nymphula) female laying eggs on aquatic plants in a pond, Wiltshire, UK, June.
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Nick Upton / Large red damselfly (Pyrrhosoma nymphula) female laying eggs on aquatic plants in a pond, Wiltshire, UK, June.
Great pond snail (Lymnaea stagnalis) grazing aquatic vegetation in a garden pond, Wiltshire, UK, February.
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Nick Upton / Great pond snail (Lymnaea stagnalis) grazing aquatic vegetation in a garden pond, Wiltshire, UK, February.
Great pond snail (Lymnaea stagnalis) grazing aquatic vegetation in a garden pond, Wiltshire, UK, February.
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Nick Upton / Great pond snail (Lymnaea stagnalis) grazing aquatic vegetation in a garden pond, Wiltshire, UK, February.
Great pond snail (Lymnaea stagnalis) grazing aquatic vegetation in a garden pond, Wiltshire, UK, May.
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Nick Upton / Great pond snail (Lymnaea stagnalis) grazing aquatic vegetation in a garden pond, Wiltshire, UK, May.
Greylag goose (Anser anser) emerging from a lake with its brood of six goslings following, Gloucestershire, UK, April.
01630406
Nick Upton / Greylag goose (Anser anser) emerging from a lake with its brood of six goslings following, Gloucestershire, UK, April.
Fulmar (Fulmaris glacialis) in flight, head on view, Cornwall, UK, April.
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Nick Upton / Fulmar (Fulmaris glacialis) in flight, head on view, Cornwall, UK, April.
Black-headed gulls (Chroicocephalus ridibundus) in fight over nesting territory on a lake margin, Gloucestershire, UK, April.
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Nick Upton / Black-headed gulls (Chroicocephalus ridibundus) in fight over nesting territory on a lake margin, Gloucestershire, UK, April.
Black-headed gull (Chroicocephalus ridibundus) male regurgitating food for its mate on their teritory on the margins of a lake, Gloucestershire, UK, April.
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Nick Upton / Black-headed gull (Chroicocephalus ridibundus) male regurgitating food for its mate on their teritory on the margins of a lake, Gloucestershire, UK, April.
Rook (Corvus frugilegus) with leatherjacket (Tipulidae) extracted from the ground by probing coastal grassland with its beak, Cornwall, UK, April.
01630402
Nick Upton / Rook (Corvus frugilegus) with leatherjacket (Tipulidae) extracted from the ground by probing coastal grassland with its beak, Cornwall, UK, April.
Rock pipit (Anthus petrosus) perched on cliff edge grassland with the sea in the background, Cornwall, UK, April.
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Nick Upton / Rock pipit (Anthus petrosus) perched on cliff edge grassland with the sea in the background, Cornwall, UK, April.
Raven (Corvus corax) perched on fence post among flowering Common gorse bushes (Ulex europaeus) on a clifftop with the sea in the background, Cornwall, UK, April.
01630400
Nick Upton / Raven (Corvus corax) perched on fence post among flowering Common gorse bushes (Ulex europaeus) on a clifftop with the sea in the background, Cornwall, UK, April.
Raven (Corvus corax) perched on fence post among flowering Common gorse bushes (Ulex europaeus) on a clifftop with the sea in the background, Cornwall, UK, April.
01630399
Nick Upton / Raven (Corvus corax) perched on fence post among flowering Common gorse bushes (Ulex europaeus) on a clifftop with the sea in the background, Cornwall, UK, April.
Group of young Mallards (Anas platyrhynchos) swimming together on the River Otter, near Otterton, Devon, UK, June.
01630398
Nick Upton / Group of young Mallards (Anas platyrhynchos) swimming together on the River Otter, near Otterton, Devon, UK, June.
Little ringed plover (Charadrius dubius) foraging for invertebrate food on the muddy margins of a freshwater lake, Gloucestershire, UK, April.
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Nick Upton / Little ringed plover (Charadrius dubius) foraging for invertebrate food on the muddy margins of a freshwater lake, Gloucestershire, UK, April.
Kestrel (Falco tinnunculus) pair copulating on cliff ledge, Cornwall, UK, April.
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Nick Upton / Kestrel (Falco tinnunculus) pair copulating on cliff ledge, Cornwall, UK, April.
Kestrel (Falco tinnunculus) male perched on a rock cairn on a coastal headland, Cornwall, UK, April.
01630395
Nick Upton / Kestrel (Falco tinnunculus) male perched on a rock cairn on a coastal headland, Cornwall, UK, April.
Marbled white butterfly (Melanargia galathea) nectaring on a Greater knapweed flower (Centaurea scabiosa) in a chalk grassland meadow, Wiltshire, UK, June.
01630520
Nick Upton / Marbled white butterfly (Melanargia galathea) nectaring on a Greater knapweed flower (Centaurea scabiosa) in a chalk grassland meadow, Wiltshire, UK, June.
Colony of Large knapweed aphids (Uroleucon jaceae) feeding on Black / Common knapweed (Centaurea nigra) stem, Mellis Common, Suffolk, UK May.
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Nick Upton / Colony of Large knapweed aphids (Uroleucon jaceae) feeding on Black / Common knapweed (Centaurea nigra) stem, Mellis Common, Suffolk, UK May.
Hairy-footed flower bee (Anthophora plumipes) female returning to her nest burrow in an earth bank to provision it with pollen attached to her hind legs, Cornwall, UK, April.
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Nick Upton / Hairy-footed flower bee (Anthophora plumipes) female returning to her nest burrow in an earth bank to provision it with pollen attached to her hind legs, Cornwall, UK, April.
Male Hairy-footed flower bee (Anthophora plumipes) wiating to intercept females nesting in an earth bank, Cornwall, UK, April.
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Nick Upton / Male Hairy-footed flower bee (Anthophora plumipes) wiating to intercept females nesting in an earth bank, Cornwall, UK, April.
Common grass-veneer moth / Crambid snout moth (Agriphila tristella) resting on grass grass flowers by day in a chalk grassland meadow, Wiltshire, UK, June.
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Nick Upton / Common grass-veneer moth / Crambid snout moth (Agriphila tristella) resting on grass grass flowers by day in a chalk grassland meadow, Wiltshire, UK, June.
Head-on view of a Goldenrod crab spider (Misumena vatia) juvenile well camouflaged on a Common hogweed (Heracleum sphondylium) flowerhead, Wiltshire, UK, June.
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Nick Upton / Head-on view of a Goldenrod crab spider (Misumena vatia) juvenile well camouflaged on a Common hogweed (Heracleum sphondylium) flowerhead, Wiltshire, UK, June.
Flat periwinkle (Littorina obtusata / littoralis) yellow form crawling over rocks exposed at low tide, Cornwall, UK, March.
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Nick Upton / Flat periwinkle (Littorina obtusata / littoralis) yellow form crawling over rocks exposed at low tide, Cornwall, UK, March.
Face fly / Autumn house fly (Musca autumnalis) dusted with pollen feeding on a Forsythia flower, Wiltshire garden, UK, March.
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Nick Upton / Face fly / Autumn house fly (Musca autumnalis) dusted with pollen feeding on a Forsythia flower, Wiltshire garden, UK, March.
Dark green fritillary butterfly (Argynnis aglaja) nectaring on a Greater knapweed flower (Centaurea scabiosa) in a chalk grassland meadow, Wiltshire, UK, June.
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Nick Upton / Dark green fritillary butterfly (Argynnis aglaja) nectaring on a Greater knapweed flower (Centaurea scabiosa) in a chalk grassland meadow, Wiltshire, UK, June.
Two Great crested newt (Triturus cristatus) males courting a female, with one arching his back and waving his tail as the other grasps her in a garden pond at night, near Wells, Somerset, UK, March. P...
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Nick Upton / Two Great crested newt (Triturus cristatus) males courting a female, with one arching his back and waving his tail as the other grasps her in a garden pond at night, near Wells, Somerset, UK, March. P...