Daniel Heuclin / Sidewinder or Horned rattlesnake (Crotalus cerastes) tracks on sand, Mojave desert, California, June.

Daniel Heuclin / Sidewinder or Horned rattlesnake (Crotalus cerastes) moving across sand, Mojave desert, California, June.

David Woodfall / Footpath and cycleway signs during January 2014 flooding, Langport, Somerset, UK, England, UK, 11th January 2014.

David Woodfall / Footpath and cycleway signs during January 2014 flooding, Langport, Somerset, UK, England, UK, 11th January 2014.

David Woodfall / Car park with footpath signs, during January 2014 floods, Langport, Somerset, UK, England, UK, 11th January 2014.

David Woodfall / Car park with footpath signs, during January 2014 floods, Langport, Somerset, UK, England, UK, 11th January 2014.

Ernie Janes / Hiker on a path looking over Muker Village, Swaledale, Yorkshire Dales National Park, Yorkshire, England, UK, November 2011

Paul Hobson / Path through autumnal Beech trees (Fagus sylvatica) near Wyming brook, Sheffield, England, October 2012.

Paul Hobson / Path through autumnal Beech trees (Fagus sylvatica) near Wyming brook, Sheffield, England, October 2012.

Gary K. Smith / Country lane lined with Cow parsley (Anthriscus sylvestris) in arable farmland, Norfolk, England, June.

Christophe Courteau / Pack of African wild dogs (Lycaon, pictus) smelling a scent on a trail, Mala Mala Game Reserve, South Africa, June.

John Waters / A half-flooded tractor track heading towards inundated farmland on West Sedgemoor near Stoke St Gregory. Somerset Levels, Somerset, UK. January 2014

Peter Lewis / Rural road through the Poppy (Papaver rhoeas) fields. South Downs National Park, Arundel, West Sussex, England, UK. June.

Roger Powell / Cart tracks in a flowering meadow, with Cork oaks (Quercus suber) in the background, Beja, Portugal, April.

Tom Gilks / Wildlife poaching patrol unit on horseback inspect wooden stakes used by poachers to impale game along trails, Mount Kenya NP, Kenya

Kirkendall-Spring / Camp site on Skyline Ridge in the Mount Baker Wilderness, Baker-Snoqualmie National Forest. Washington, USA, August 2013.

Tony Phelps / Leopard tortoise (Stigmachelys pardalis) crossing a road with a car approaching, Swartberg Mountains, Western Cape, South Africa.

Andy Rouse / Cheetah (Acinonyx jubatus) cubs walking along track, Maasai Mara, Kenya, Africa

Adrian Davies / Rhododendron (Rhododendron x. superponticum) invasive species, growing next to pathway, Snowdonia National Park, North Wales Wales

Adrian Davies / Rhododendron (Rhododendron x. superponticum) growing on slopes of mountain. Snowdonia National Park, North Wales Wales

Adrian Davies / Rhododendron (Rhododendron x. superponticum) flowering on mountain footpath, invasive species, Snowdonia National Park, North Wales Wales

Adrian Davies / Rhododendron (Rhododendron x. superponticum) flowering, invasive species, Snowdonia National Park, North Wales Wales

Floris van Breugel / Moonrise over the flooded Alvord Desert with photographer Floris van Breugel watching the sky. Long exposure. Harney County, Oregon. January 2010

Roland Seitre / Kagus (Rhynochetos jubatus) with ranger repairing track, Parc Provincial de la Riviere Bleue / Blue River Provincial Park, Yate, South Province, New Caledonia, August 2012. Endangered species

Wild Wonders of Europe / Möllers / Wooden signposts for hiking trails in Tarcu Mountains Natura 2000 site at the Meteorological Station of Cuntu. Southern Carpathians, Muntii Tarcu, Caras-Severin, Romania, October 2012

Andrew Parkinson / Gannet (Morus bassanus) colony, cliff top view showing the trails that gannets leave during a 2 second exposure. Shetland Islands, Scotland, UK, September

Andrew Parkinson / Gannet (Morus bassanus) colony, cliff top view showing the trails that gannets leave during a 2 second exposure. Shetland Islands, Scotland, UK, September

Andrew Parkinson / Gannet (Morus bassanus) colony, cliff top view showing the trails that gannets leave during a 2 second exposure. Shetland Islands, Scotland, UK, September

Andrew Parkinson / Gannet (Morus bassanus) colony, cliff top view showing the trails that gannets leave during a 2 second exposure. Shetland Islands, Scotland, UK, September

Andrew Parkinson / Gannet (Morus bassanus) colony, cliff top view showing the trails that gannets leave during a 2 second exposure. Shetland Islands, Scotland, UK, September

Yukihiro Fukuda / Japanese fireflies (Luciola cruciata) in flight at night, endemic species, Yaku-shima UNESCO World Heritage Site, Kagoshima, Japan, May

Pascal Tordeux / A path from the peaks in Vars Queyras massif, above the park Escreins, France, August 2011

Bence Mate / Star trails, 20 minutes exposure time, Pusztaszer, Hungary

Andy Rouse / Aerial view of Lesser Flamingo (Phoenicopterus / Phoeniconaias minor) walking through shallow water, leaving trails in the alkaline mud. Lake Natron, Tanzania, September.

Inaki Relanzon / Joshua tree (Yucca brevifolia) at night, with star trails, Joshua Tree National Park, California, USA, June 2012.

Juergen Freund / Night over Mungerannie Wetland with a red dune over Derwent River, South Australia

Kirkendall-Spring / Elephant Canyon at night with star trails. The Needles District of Canyonlands National Park, Utah, October 2012.

George Sanker / Appalachain Trail running through forest in Baxter State Park, Maine, USA. October 2012.

Kirkendall-Spring / Camping tent at night in Sawtooth Lake area with Mount Regan on the skyline in the Sawtooth Wilderness. Idaho, USA, July 2011

Kirkendall-Spring / Sawtooth Lake and Mount Regan in the Sawtooth Wilderness as seen form Trail #640. Idaho, USA, July 2011

Kirkendall-Spring / Hiker on the Boundary Trail with McGowin peak in the background - Sawtooth Wilderness. Idaho, USA, July 2011. Model released

Kirkendall-Spring / Hiker on the Boundary Trail with McGowin Peak in the background - Sawtooth Wilderness. Idaho, USA, July 2011. Model released

Kirkendall-Spring / Hiker's feet in boots walking across the Plains of Abraham on the Loowit Trail #216 in Mount St Helens National Volcanic Monument. Washington, USA, August 2011.

Kirkendall-Spring / Hiker's legs and boots crossing the Plains of Abraham on the Loowit Trail #216 in Mount St Helens National Volcanic Monument. Washington, USA, August 2011. Model released.

Kirkendall-Spring / Man hiking along the Boundary Trail #1 below Norway Pass in Mount St Helens National Volcanic Monument, Washington, USA. August 2011. Model released.

Kirkendall-Spring / Boundary Trail #1 below Norway Pass in Mount St Helens National Volcanic Monument. Washington, USA, August 2011.

Gavin Hellier / Blurred motion light trails in an train tunnel under the Huangpu river linking the Bund to Pudong, Shanghai, China 2010

Gavin Hellier / Pasadena Freeway, CA Highway 110, at dusk with light trails from cars, leading into downtown Los Angeles, California, USA, June 2011

Juan Carlos Munoz / Aurora borealis and star trails over Jokulsarlon glacier lagoon. Southern Iceland, Europe, August 2012.

Niall Benvie / Aeroplane flying overhead, leaving contrails / condensation-trails against blue sky. Scotland, March.

Kirkendall-Spring / Man hiking along the Death Canyon Shelf on the Teton Crest Trail in Grand Teton National Park. Wyoming, USA, July 2011

Kirkendall-Spring / Night time view of a campsite tent on the Death Canyon Shelf along the Teton Crest Trail in Grand Teton National Park. Wyoming, USA, July 2011

Kirkendall-Spring / Night time view of a campsite tent on the Death Canyon Shelf along the Teton Crest Trail in Grand Teton National Park. Wyoming, USA, July 2011

Kirkendall-Spring / Bridge crossing Daley Bay along the Kab-Ash Trail in Voyageurs National Park. Minnesota, USA, August 2011

Kirkendall-Spring / Hiker on the Silver Trail / North Country Trail in Jay Cook State Park. Minnesota, USA, August 2011 Model released

Kirkendall-Spring / Hiker on the Carlton Trail in Jay Cook State Park. Minnesota, USA, August 2011. Model released

Wild Wonders of Europe / Widstrand / Park ranger with pack mules on the main trail, Paklenica National Park, Velebit Nature Park, Rewilding Europe area, Velebit mountains, Croatia June 2012

Alex Hyde / Man and woman walking along track, rear view, above Monsal Dale near Brushfield, Peak District National Park, Derbyshire, UK. August 2012

Alex Hyde / Hiker in Vallone di Valelle, Gran Paradiso National Park, Aosta Valley, Pennine Alps, Italy. July 2012

Alex Hyde / Vallone di Valelle, Gran Paradiso National Park, Aosta Valley, Pennine Alps, Italy. July.

Alex Hyde / Hiker in Vallone di Valelle, Gran Paradiso National Park, Aosta Valley, Pennine Alps, Italy. July.

Alex Hyde / Path leading towards Rifugio Ferraro, Resy, Aosta Valley, Monte Rosa Massif, Pennine Alps, Italy. July 2012

Floris van Breugel / A wide variety of fluorescent minerals including scheelite (blue/white), calcite (red), silica coatings containing uranyl ions (green), fluorite (pink), and 'desert varnish' (yellow/orange), populate...

Oriol Alamany / Hiker walking the GR11 long distance footpath near Queralbs in Ribes valley, Pyrenees, Ripollas, Catalonia, Spain (Model released)

Oriol Alamany / Hiker walking the GR11 long distance path, Albera Natural Park, Pyrenees, Alt Emporda, Catalonia, Spain

Christophe Courteau / Aerial View showing Hippopotamus' trails in the swamps of the Okavango Delta, Botswana

Christophe Courteau / Aerial view of Okavango delta game trails (hippos) lead to waterhole Botswana

Christophe Courteau / Aerial view of Okavango Delta, Botswana, Southern-Africa. End of rainy season, April.

Christophe Courteau / Aerial view of African elephant herd {Loxodonta africana}. Okavango delta, Botswana

John Waters / Animal trail traversing grass clearing in small copse. Gower Peninsula, Wales, UK, November 2011.

Peter Cairns / 2020VISION / Gannet (Morus bassanus) breeding colony, long exposure showing flight trails. Hermaness National Nature Reserve, Shetland, Scotland, July.

Eric Medard / Whiskered Bat (Myotis mystacinus) drinking from water in flight, with star-trails in night sky. France, Europe, August.

Mark Hamblin / 2020VISION / Walker on woodland trail through pinewoods, Beinn Eighe National Nature Reserve, Scotland, UK, model released, June 2011

Mark Hamblin / 2020VISION / Trail through Abernethy Forest NNR. Cairngorms National Park, Scotland, UK, June 2011.

Billy Black / Long exposure of car lights passing a tree decorated with lights, Thailand. March 2006.

Richard Du Toit / Dry waterhole with game trails, Western Cape, South Africa

Nick Turner / South West Coast Path National Trail waymarker information sign, near Durdle Door, Dorset, UK, May 2009.

Charlie Summers / Long-exposure night photography reveals the rotation of the Earth, producing star trails centred on the Northern Circumpolar Star, Polaris. Toadstool State Park, Nebraska.

Charlie Summers / Long-exposure night photography reveals the rotation of the Earth, producing star trails centred on the Northern Circumpolar Star, Polaris. Toadstool State Park, Nebraska.

Nick Upton / 2020VISION / White-lipped snail (Cepaea hortensis) crawling over Ivy leaf (Hedera helix) in garden, with house in background, Wiltshire, England, UK, October . Property released. Did you know? Snails sometimes mov...

Rob Jordan / 2020VISION / Hikers on the Red Kite trail in the Urban Red Kite area of the Derwent Valley, Gateshead, Tyne and Wear, UK, on the edge of Tyneside following on from the 'Northern Kites' re-introduction programme be...

Rob Jordan / 2020VISION / Red Kite trail information sign in the Urban Red Kite area of the Derwent Valley, Gateshead, Tyne and Wear, UK, on the edge of Tyneside following on from the 'Northern Kites' re-introduction programme...

Rob Jordan / 2020VISION / Red Kite trail sign in the Urban Red Kite area of the Derwent Valley, Gateshead, Tyne and Wear, UK, on the edge of Tyneside following on from the 'Northern Kites' re-introduction programme between 200...

Rob Jordan / 2020VISION / Red Kite trail map in the Urban Red Kite area of the Derwent Valley, Gateshead, Tyne and Wear, UK, on the edge of Tyneside following on from the 'Northern Kites' re-introduction programme between 2004...

Rob Jordan / 2020VISION / Red Kite trail information board in the Urban Red Kite area of the Derwent Valley, Gateshead, Tyne and Wear, UK, on the edge of Tyneside following on from the 'Northern Kites' re-introduction programm...

Rob Jordan / 2020VISION / Red Kite Trail sign in the Urban Red Kite area of the Derwent Valley, Gateshead, Tyne and Wear, UK, on the edge of Tyneside following on from the 'Northern Kites' re-introduction programme between 200...

Rob Jordan / 2020VISION / Hikers read information board on the Red Kite trail in the Urban Red Kite area of the Derwent Valley, Gateshead, Tyne and Wear, UK, on the edge of Tyneside following on from the 'Northern Kites' re-in...

Kirkendall-Spring / Person hiking along the Burroughs Loop Trail near Sunrise in Mount Rainier National Park, Washington, USA, August 2009

Stefan Huwiler / Matterhorn at sunrise in autumn with reflection in the Riffelsee and star-trails. Zermatt, Valais, Switzerland, Europe, October 2008.

Charlie Summers / Star trails above Colorado. USA, April.

Charlie Summers / Star trails in a South Dakota sky over a Ogallala Lakota Sioux teepee. USA, April.

Adrian Davies / Grazing trails left by Common / Garden Snail (Helix aspersa) as they fed on algea on a post. UK, April.

Adrian Davies / Grazing trails left by Common / Garden Snail (Helix aspersa) as they fed on algea on a post. UK, April.

Yva Momatiuk & John Eastcott / Scribbly Gum (Eucalyptus haemastoma) with moth trails in bark, New South Wales, Australia

Inaki Relanzon / Thousand-year-old Sabina / Canary Juniper, (Juniperus turbinata canariensis) against a night sky with star trails. El Sabinar, El Hierro Island, Canary Islands, , February 2011.