John Cancalosi / Red deer stag (Cervus elaphas) in rut, calling. UK, Europe

John Cancalosi / Red deer stag (Cervus elaphus) calling during rut. England, UK, Europe

Edwin Giesbers / Red deer (Cervus elaphus) silhouetted stag bellowing during rut in grassland, doe in background. Richmond Park, London, England, UK. October

Andy Rouse / Bengal tiger (Panthera tigris) with mouth open, cooling in pool during summer heat. Ranthambhore National Park, India.

Loic Poidevin / Lion (Panthera leo) roaring, male, Masai Mara, Kenya.

Oscar Dewhurst / Red Deer (Cervus elaphus) stag roaring silhouetted in grassland at sunrise. Richmond Park, London. September

Oscar Dewhurst / Red deer (Cervus elaphus) stag silhouetted, roaring in the mist before sunrise. Richmond Park, London. September

Oscar Dewhurst / Red deer (Cervus elaphus) stag roaring in the mist, silhouetted, backlit at sunrise. Richmond Park, London, UK. October

Oscar Dewhurst / Red deer (Cervus elaphus) stag roaring with bracken-covered antlers. Richmond Park, London, UK. October

Oscar Dewhurst / Red deer (Cervus elaphus) roaring in the bracken in the mist at sunrise. Bushy Park, London, UK. September. Cropped

Oscar Dewhurst / Red deer (Cervus elaphus) stag in misty conditions bellowing whilst sat down in long grass with the rising sun in the background. Richmond Park, London, UK. September

Oscar Dewhurst / Red deer (Cervus elaphus) stag roaring in long grass. Richmond Park, London, UK. October